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Umbrella cockatoo (Cacatua alba), portrait. Captive
Seeds and seedpods of Armenian grape hyacinth (Muscari armeniacum) June
Grey whale (Eschrichtius robustus) eye, Magdalena Bay, Baja California, Mexico, February. Winner in the Ocean Views category of the 2018 Natures Best Winland Smith Rice Awards
Polar bear (Ursus maritimus) walking on Champ Island glacier above the sea, Franz Josef Land, Russian Arctic
Bobcat (Lynx rufus) standing on branch in snow. Madison River Valley, Yellowstone National Park, Wyoming, USA. January
Green turtle (Chelonia mydas) with rays of sunlight, black and white image, Akumal, Caribbean Sea, Mexico, July. Second place in the Visions of our Nature 2018
English springer spaniel, three standing with front legs on rock, Rhododendron flowers in background. Haddam, Connecticut, USA. June
Grey whale (Eschrichtius robustus) tail, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Living stone plant (Lithops dorotheae) one of the living stones plants, in cultivation. It is in transitional stage, with the old leaves withering and new ones emerging. Native to South Africa
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
English oak tree (Quercus robur) in field, with early spring growth, Gloucestershire, UK, May
Bichon frise beside Colorado blue columbine (Aquilegia coerulea) flowers. Old Lyme, Connecticut, USA. May
Bichon frise standing on rock amongst Rhododendron flowers. Haddam, Connecticut, USA. May
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Seedling cone plants (Conophytum subfenestratum), approximately two months after sowing. Focus-stacked image
Fork-leaved sundew (Drosera bipinata) carnivorous plant, native to Australia
White witch moth (Thysania agrippina) camouflaged against bark of tree. Manu Biosphere Reserve, Amazonia, Peru
Newfoundland dog, male standing in snow. Killingworth, Connecticut, USA
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
RF - Maidens veil / Bridal veil fungus (Phallus indusiatus) with indusium fully formed, on rainforest floor. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Red fox (Vulpes vulpes) visiting woodland stream to drink at night. Camera trap image. Near Bath, England, UK. June
Tufted / Brown capuchin (Cebus apella), female with baby on back, sitting in tree, mid-altitude montane forest, Manu Biosphere Reserve, Peru
Rainbow at Iguazu Falls, Brazil / Argentina border. Photographed from Argentina. August 2017
Princely fiddler crab (Uca princeps), Bahia Magdalena, Baja California Peninsula, Mexico, June
Tourist touching Grey whale (Eschrichtius robustus) calf, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, February
Grey whale (Eschrichtius robustus) breaching, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Caribbean flamingo (Phoenicopterus ruber) walking, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September
Birch tree (Betula sp) next to waterfall, Iceland. Runner-Up in the Plants category of the International OasisPhotoContest 2018
English springer spaniel in Rhododendron. Haddam, Middlesex, Connecticut, USA
Weimaraner in front of Rhododendron flowers, Haddam, Connecticut, USA. May
Parson's chameleon (Calumma parsonii)Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
Green-veined white butterfly (Pieris napi) Whitelye Common Nature Reserve, Monmouthshire, Wales, UK
Bronzed frog / Common Wood Frog (Hylarana temporalis) Sri Lanka
Cheiridopsis species, seed capsule open in response to rain. Namaqualand, South Africa
Bengal tiger (Panthera tigris tigris) in forest clearing. Camera trap image. Pench National Park, Madhya Pradesh, India. January 2018
English springer spaniel standing in snow covered grassland, shore of Long Island Sound, Guilford, Connecticut, USA
English springer spaniel standing in front of Rhododendron flowers. Haddam, Connecticut, USA. June
Black skimmer (Rynchops niger) foraging along surf line. Tierra Verde, Florida, USA
Chocolate labrador retriever, two puppiessitting in spring flowers. Haddam, Connecticut, USA. June
Shetland sheepdog running in Common bluet (Houstonia caerulea) field. Haddam, Connecticut, USA. May
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Bush cricket or katydid (Tettigoniidae) with spines, camouflaged amongst moss. Forest interior of mid-altitude montane rainforest, Manu Biosphere Reserve, Peru
Hoatzin (Opisthocomus hoazin), pair on floating log in Cocha Salvador ox-bow lake. Manu Biosphere Reserve, Peru
RF - Red-and-green macaw (Ara chloropterus) flock feeding and flying. Wall of clay lick, Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Aerial view of lowland Amazonia rainforest in morning mist. Manu Biosphere Reserve, Amazonia, Peru. November 2017
Grey whale (Eschrichtius robustus) calf, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Red fox (Vulpes vulpes) cub exploring woodland. Near Bath, England, UK. May
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Pink-toed tarantula (Avicularia avicularia), female waiting in ambush outside daytime lair in palm tree trunk. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Bell glass frog (Cochranella nola) from below, photographed on a pane of glass in Manu Biosphere Reserve, Peru
Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017
Marsh deer (Blastocerus dichotomus), stag. Pousada Araras, northern Pantanal, Mato Grosso, Brazil
Red ruffed lemur (Varecia rubra) female in understorey of lowland rainforest. Masoala National Park, north-east Madagascar
Giant Anteater (Myrmecophaga tridactyla), two sniffing air in savanna. Caiman Ecological Refuge, Southern Pantanal, Moto Grosso do Sul, Brazil
Silky sifaka (Propithecus candidus), female with baby sitting in rainforest understorey. Mid-altitude montane rainforest, Marojejy National Park, north-east Madagascar
Saltpan, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, February
Snowy Egret (Egretta thula), Guerrero Negro, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Western Grebe (Aechmophorus occidentalis) swimming, Guerrero Negro, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Blurred motion image of Caribbean flamingo (Phoenicopterus ruber) juveniles, captured to be tagged and released to the wild, captive, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico
Finespotted Jawfish (Opistognathus punctatus), Bahia Magdalena, Baja California Peninsula, Mexico, June
Dune and Red Mangrove (Rhizophora mangle), Bahia Magdalena, Baja California Peninsula, Mexico, June
Long Beaked Common Dolphin (Delphinus capensis), Espiritu Santo National Park, Sea of Cortez (Gulf of California), Mexico, February
Red Mangrove (Rhizophora mangle) seedling, Bahia Magdalena, Baja California Peninsula, Mexico, June
Bottlenose Dolphin (Tursiops truncatus) in mangrove canals, Bahia Magdalena, Baja California Peninsula, June
Grey whale (Eschrichtius robustus), Magdalena Bay, Baja California, Mexico, February
Caribbean Flamingo (Phoenicopterus ruber) juvenile, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September
European hare (Lepus europaeus), two leverets crouched in soil. Navarra, Spain. August
Black-necked Stilt (Himantopus mexicanus) walking, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September
Pyrenean squill (Scilla lilio-hyacinthus), Ibarra, Orozko, Biscay, Basque Country, Spain. March
Sunflower (Helianthus annuus), close-up. Sunflower plantation in Cuestahedo, Merindad de Montija, Merindades, Burgos, Castilla y Leon, Spain. August
Iberian ibex (Capra pyrenaica) amongst scrub, Cliffs of Maro-Cerro Gordo Natural Area, Granada, Andalusia, Spain. January
White-tailed deer (Odocoileus virginianus), leucistic white doe, New York, USA, June
Griffon vulture (Gyps fulvus) landing amongst other vultures on cliff above sea, Liendo Valley within the Montana Oriental Costera, Cantabria, Spain
RF - Amazon river dolphin (Inia geoffrensis) at surface, Rio Negro, Manaus, Brazil
Hippopotamus (Hippopotamus amphibius) in pool with mouth open. Mana Pools National Park, Zimbabwe
Andean cock-of-the-rock (Rupicola peruvianus), male at a lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Bengal tiger (Panthera tigris tigris), male near tree previously spray marked by female. Camera trap image. Pench National Park, Madhya Pradesh, India. January 2018
Grey whale (Eschrichtius robustus) tail, Magdalena Bay, Baja California, Mexico, February
Red-and-green macaw (Ara chloropterus) flock flying in front of clay lick. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Jaguar (Panthera onca palustris) female yawning with mouth wide open, wearing Oncafari Project radio collar. Caiman Lodge, southern Pantanal, Mato Grosso do Sul, Brazil. September 2017
Caribbean flamingo (Phoenicopterus ruber) abandoned egg in nest at the end of the breeding season, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September
Grey whale (Eschrichtius robustus) calf, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, February
Caribbean flamingo (Phoenicopterus ruber) juvenile begging, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September