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View towards Naranjo de Bulnes (Picu Urriellu) a distinctive 2519m limestone peak, Picos de Europa mountains, viewed from Arenas de Cabrales at sunset, Asturias, Spain, August 2016
Bolivian red howler (Alouatta sara), Pampas del Yacuma Protected Area, Bolivia
Pyrenean merendera / False meadow saffron (Merendera pyrenaica / Colchicum montanum) flowering on montane pastureland, Vegas de Sotres, Picos de Europa, Asturias, Spain, August
Mediterranean sea holly / Pyrenean eryngo (Eryngium bourgatii) flowering on montane pastureland, Covadonga, Picos de Europa, Asturias, Spain, August
Cornish heath (Erica vagans) clump flowering on montane pastureland, above the Lakes of Covadonga, at 1300m, Picos de Europa, Asturias, Spain, August
Aerial view Fin whales (Balaenoptera physalus) lunge-feeding in the southern Sea of Cortez (Gulf of California), Baja California, Mexico
White rhinoceros (Ceratotherium simum) male and female, face to face, Mosi Oa Tunya National Park, Zambia
Black rhinoceros (Diceros bicornis) stands in evening light on Chiefas Island in Okavango DeltaBlack rhinoceros (Diceros bicornis) stands in evening light on Chiefs Island in Okavango Delta, Botswana, . This rhino was released in the Okavango Delta as part of efforts to rebuild the rhino
Stoat (Mustela erminea) Creux du Van, Jura Suisse, Switzerland, September
White tent making bat (Ectophylla alba) roosting in Heleconia leaf, La Selva Field Station, Costa Rica
Gorner glacier with Mont Rose and Lyskamm, Swiss Alps, Valais, Switzerland. September 2018
Olive-backed sunbird (Nectarinia jugularis) female returning to her nest with nesting material, Cape Hillsborough National Park, Queensland, Australia, September
Emu (Dromaius novaehollandiae) juvenile, Cleland Wildlife Park, South Australia, March. Captive
Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive
Silhouette of an Agile wallaby (Macropus agilis) hopping along the beach at dusk, Cape Hillsborough National Park, Queensland, Australia. September
Rainbow lorikeet (Trichoglossus moluccanus) on a branch, Cania Gorge National Park, Queensland, Australia. September
An Orange lacewing butterfly (Cethosia penthesilea), Cairns Botanical Gardens, Queensland, Australia. Captive. August
A Perentie (Varanus giganteus) using its forked tongue to detect smells, Alice Springs Reptile Centre, Northern Territory, Australia, Captive. May
Orca (Orcinus orca) surfacing and blowing, Shetland, Scotland, UK, August
Otter (Lutra lutra) portrait on shore with boats in background. Argyll, Scotland, UK, August
RF - Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August
RF - Sand dunes with Marram grass (Ammophila arenaria) and beach at Seilebost beach, Isle of Harris, Scotland, UK. October
Banded demoiselle (Calopteryx splendens) male, resting on grass head, Lower Tamar Lakes, Cornwall, UK. June
American oystercatcher (Haematopus palliatus) adult feeding chick, Long Island, New York, USA. June
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Common loons (Gavia immer), two adults and chick swimming on a misty lake in early morning, Michigan, USA. June
Common redpoll (Carduelis flammea), male perched in alder (Alnus rugosa) during a snowstorm in winter, New York, USA, January
Western Grebe (Aechmorphus occidentalis), pair during rushing courtship display in which they run across waters surface in synchrony, near Escondido, California, USA, January
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Horned Puffin (Fratercula corniculata) flying over a pattern of kelp and sea foam, photographed from above, St. Paul, Pribilof Islands, Alaska, USA. July
Roseate spoonbill (Ajaia ajaja) adult in breeding plumage preening tail feathers, Black Point Wildlife Drive, Merritt Island National Wildlife Refuge, Florida, USA, January
Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January
Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January
Bengal tiger (Panthera tigris tigris) walking at night dominant male (T29) with monsoon clouds and lightning. Kanha National Park, Central India
Bengal tiger (Panthera tigris tigris) dominant male (T29) cooling off in a waterhole Kanha National Park, Central India. Camera trap image
Bengal tiger (Panthera tigris tigris) dominant male (T29) walking on mud dam wall. Kanha National Park, Central India. Camera trap image
Bengal tiger (Panthera tigris tigris) tigress (T32) feeding on Spotted deer / Chital (Axis axis) killed by her earlier that day while her young cub watches. Kanha National Park, Central India
Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image
RF - Brown hare, (Lepus europaeus) male pursuing female that is in season, Islay, Scotland, UK. March (This image may be licensed either as rights managed or royalty free.)
RF - Brown hare, (Lepus europaeus), male and female displaying courtship behaviour, Islay, Scotland, UK. March (This image may be licensed either as rights managed or royalty free.)
RF - Black Grouse (Tetrao tetrix), two males fighting on lek, Scotland, UK. AprilRF - Black Grouse (Tetrao tetrix), two males fighting on lek, Scotland, UK.April (This image may be licensed either as rights managed or royalty free.)
Red squirrel, (Sciurus vulgaris), silhouetted on stump at dawn, Scotland, UK. MarchRed squirrel, (Sciurus vulgaris), silhouetted on stump at dawn, Scotland, UK.March
Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February