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Harvest mouse (Micromys minutus) climbing among Cow Parsley, Hertfordshire, England, UK, May, Controlled conditions
Magpie inkcap (Coprinopsis picacea) uncommon inkcap that usually grows singularly often under beech trees, Bedfordshire, England, UK, October. Focus stacked image
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Caribbean Flamingo (Phoenicopterus ruber) chick under the wing of protective parent, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Sunrays in Cenote Jardin del Eden, Quintana Roo, Yucatan Peninsula, Mexico, May 2016. Photographed for The Freshwater Project
Aerial shot of the Laguna Sian Ka an, freshwater lagoon, at edge of the sea, close to Boca Paila, Quintana Roo, Yucatan Peninsula, Mexico. Photographed for The Freshwater Project February 2014
Rainbow river, Rainbow Springs State Park, Florida, USA. January. Photographed for The Freshwater Project
Fairy inkcap fungus (Coprinellus disseminatus) small group of this inkcap growing from mossy log, Oxfordshire, England, UK, October. Focus Stacked Image
Caribbean Flamingo (Phoenicopterus ruber) chick group walking around the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Pine marten (Martes martes) reflected in water, Ardnamurchan Peninsula, west coast of Scotland, UK
Pod of Risso Dolphin (Grampus griseus) at surface, Baja California, Mexico
European red fox (Vulpes vulpes crucigera) in deep snow in front of steep rocks. Gran Paradiso National Park, Italy
Short finned pilot whale (Globicephala macrorhynchus) about to surface, Baja California, Mexico, February
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Humpback whale (Megaptera novaeangliae) breaching, Baja California, Mexico
Humpback whale (Megaptera novaeangliae) breaching - leaping out of the water, Sea of Cortez, Baja California, Mexico
Harvest mouse (Micromys minutus) Sitting on frosty seedhead, Hertfordshire, England, UK, January, Controlled conditions
Ivy bee (Colletes hederae) new species to the UK in 2001. Male resting on Blackberry, Oxfordshire, England, UK, September
Harvest mouse (Micromys minutus) climbing among wheat, Hertfordshire, England, UK, August, Controlled conditions
Long beaked common dolphin (Delphinus capensis) pod porpoising Baja California, Mexico
Californian Sealion (Zalophus californianus) in kelp (Macrocystis pyrifera) forest, San Benito del Este Island, Baja California Peninsula Pacific Islands Biosphere Reserve, Baja California, Mexico
Glanville fritillary butterfly (Melitaea cinxia) roosting on larval foodplant Ribwort plantain (Plantago lanceolata), UK - Captive - Focus Stacked Image
Purple hairstreak butterfly (Neozephyrus quercus) female with wings open, Hertfordshire, England, UK, July. Focus Stacked Image
Great Bustard (Otis tarda) males displaying, Salisbury Plain, Wiltshire, UK, April
Firecrest (Regulus ignicapilla) singing, Wiltshire, England, UK, April
Ants (Formicidae) attacking newly emerged cicada (Cicadacae), Yasuni National Park, Ecuador
Essex skipper butterfly (Thymelicus lineola) male visiting flower, , Wiltshire, England, UK, July
Aerial view of the Northern Rockies, Muskwa-Kechika Protected Area, British Columbia, Canada. July 2011. Photographed for The Freshwater Project
Vermilion waxcap fungi (Hygrocybe miniata) Buckinghamshire, England, UK, September - Focus Stacked Image
Amethyst deceiver toadstools (Laccaria amethystina) growing up from mossy log, Buckinghamshire, England, UK, October. Focus Stacked Image
Wood White butterfly (Leptidea sinapis) on Germander Speedwell (Veronica chamaedrys), Surrey, England, UK, May - Focus Stacked
Bullfinch (Pyrrhula pyrrhula) male walking along mossy twig, Buckinghamshire, England, UK, February
Brimstone butterfly (Goneopteryx rhamni) male roosting on Cowslip (Primula veris) Bedfordshire, England, UK, April
Bewicks swan (Cygnus columbianus) in flight with rainbow, Gloucestershire, England, UK, February
Whitethroat (Sylvia communis) male perched on Willow branch in spring, Wiltshire, England, UK, April
Sedge Warbler (Acrocephalus schoenobaenus) Greylake RSPB Reserve, Somerset Levels, England, UK, August
Orange tip butterfly (Anthocharis cardamines) female and male, visiting Honesty flower (Lunaria annua) Wiltshire, England, UK, May
Reed warbler (Acrocephalus scirpaceus) with insect larvae in beak, Greylake RSPB Reserve, Somerset Levels, England, UK, June
Great pied hoverfly (Volucella pellucens) feeding from bramble flower, Wiltshire, England, UK, July. Bumblebee mimic species
Water lilies in the Cenote Nicte Ha, Quintana Roo, Yucatan Peninsula, Mexico. May. Photographed for The Freshwater Project
Rainbow river, Rainbow Springs State Park, Florida, USA. January 2012. Photographed for The Freshwater Project
Water hyacinth (Eichhornia sp. ) and red Ludwigia (Ludwigia sp)Water hyacinth (Eichhornia sp.) and red Ludwigia (Ludwigia sp.) in a clear stream, Pantanal, Brazil. Photographed for the Freshwater Project
Lagoa Misteriosa at the bottom of a sink hole in limestone region, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil November 2016. Photographed for The Freshwater Project
Piraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp. ) Rio Olhio d AguaPiraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp.) Rio Olhio d Agua, tributary of Rio da Prata, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul
Iceberg in the sea outside the Ilulissat Icefjord UNESCO World Heritage Site, Greenland, August 2014. Photographed for the Freshwater Project
Aerial view of meltwater lake on the ice cap north-east of Sermeq Kujalleq Glacier, Ilulissat Icefjord UNESCO World Heritage Site, Sermersuaq / Greenland ice sheet, Greenland, August 2014
Aerial view of meltwater channels in ice cap north-east of Sermeq Kujalleq Glacier, Sermersuaq / Greenland ice sheet, Greenland, August 2014. Photographed for The Freshwater Project
Rift between the North American and Eurasian tectonic plates at Thingvellir, Thingvellir National Park, UNESCO World Heritage Site, Iceland, September. Photographed for the Freshwater Project
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Sunrays in Cenote Jardin del Eden, Quintana Roo, Yucatan Peninsula, Mexico. Photographed for The Freshwater Project May 2016
School of Scaled herring or Scaled sardine (Harengula clupeola), entering from the sea into the freshwater of the Laguna Yal Ku, Quintana Roo, Yucatan Peninsula, Mexico
Anhumas Abyss, a 72 meters deep cave with underwater limestone cones up to 20 meters high, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil November 2016
Aerial view of flooded forests, Anavilhanas Archipelago, Anavilhanas National Park, in the Rio Negro, Amazonas, Brazil February 2011. Photographed for The Freshwater Project
Anhumas Abyss, a 72 meters deep cave, with underwater limestone cones up to 20 meters high, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil November 2016
Candover Brook, tributary of the River Itchen, chalk stream Hampshire, England, UK, June. Photographed for The Freshwater Project
Candover Brook, tributary of the River Itchen, chalk stream Hampshire, England, UK, June, Photographed for The Freshwater Project
Split level view of the River Itchen, chalk stream at Winnall Moors, Winchester, Hampshire, England, UK, May. Photographed for The Freshwater Project
Motelon creek frozen in winter, Jogne river tributary, Saane river tributary, Gruyere, Fribourg, Switzerland, February 2012. Photographed for the Freshwater Project
Red water caused by tannins in peat bog flowing out of the Etang de la Gruere, peat bog, Rhine river catchment, Canton of Jura, Switzerland. November 2013. Photographed for the Freshwater Project
Snowy landscape with Sense River. Canton Fribourg, Switzerland, January 2001 Photographed for the Freshwater Project
Etang de la Gruere, peat bog covered in snow, Rhine river catchment, Canton of Jura, Switzerland. November 2013 Photographed for the Freshwater Project
Lac de Fenetre, alpine lake, still under an ice cover at the end of May, 2456 m above sea level, Canton of Valais, Switzerland. May 2012. Photographed for The Freshwater Project
Underwater view of the Sense river, Cantons of Fribourg and Bern, Switzerland. October 2011. Photographed for The Freshwater Project
Meltwater lake with cryoconite, on the surface of the Gorner Glacier Valais Alps, Canton Valais / Wallis, Switzerland. September. Photographed for The Freshwater Project
Frozen meltwater channels on Gorner Glacier Valais Alps, Canton Valais / Wallis, Switzerland. April 2013. Photographed for The Freshwater Project
Underwater in a cryoconite hole, a hole formed by windblown dust and particles which warm up ice of glacier and creates meltwater holes
Gorner Glacier with meltwater channels, and Breithorn, Valais Alps, Canton Valais / Wallis, Switzerland, August 2013. Stitched panorama. Photographed for The Freshwater Project
Chub (Squalius cephalus), on spawning ground Sava Bohinjka River, Slovenia June 2013. Photographed for The Freshwater Project
Rows of racks used in oyster farming at high tide, Isle de Re, Charente-Maritime, France, July 2017
Mudflat, Aiguillon Bay, Vendee, France, July 2017
Alpine chamois (Rupicapra rupicapra rupicapra) in winter landscape with snow covered trees, Valsavarenche, Gran Paradiso National Park, Italy
Goldfinch (Carduelis carduelis) perched on frost covered Teasel (Dipsacus fullonum), Hertfordshire, England, UK, January
Caribbean Flamingo (Phoenicopterus ruber) feeding three day old chick at breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, June
Caribbean Flamingo (Phoenicopterus ruber) chick creche in front of attentive adult flamingo group, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Caribbean Flamingo (Phoenicopterus ruber) tending to newborn chick while another feed two day old chick, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Caribbean Flamingo (Phoenicopterus ruber) preening chick in the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico
Aerial view of Caribbean Flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May
Harvest mouse (Micromys minutus) climbing on frosty seedhead, Hertfordshire, England, UK, January, Controlled conditions
Kingfisher (Alcedo atthis) male taking off from water after diving for fish, Worcestershire, England, UK, July
Banded demoiselle (Calopteryx splendens) male close up detail of head and thorax, Hertfordshire, England, UK, June. Captive. Focus stacked