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Western reef egret (Egretta gularis) running through shallow water with wings open, chasing small fish, Kotu creek, The Gambia
Willowherb hawkmoth (Proserpinus proserpina) resting on a leaf, Gavarnie, South of France. June
Two Clouded magpie moth (Abraxas sylvata) resting on a fern frond, Gosford Forest Park, County Armagh, Northern Ireland. June
Queen butterfly (Danaus gilippus) expanding wings after emerging from chrysalis on Aquatic milkweed (Asclepias perennis). Hill Country, Texas, USA. Sequence 12/12
Buff-necked ibis (Theristicus caudatus) with wings outstretched, Pocone, Brazil
Ventral view of Swallowtail butterfly (Papilio machaon) in flight. The Netherlands. July
Hills sheathtail bat (Taphozous hilli) flying out of an abandoned mine in late evening, Tennant Creek, Northern Territory, Australia
Eastern horseshoe bat (Rhinolophus megaphyllus) flying out from an abandoned mine in late evening. Its orange colouring is due to bleaching from the intense ammonia atmosphere in the mine
Robin (Erithacus rubecula) carrying a red berry in its beak as it takes off, Parainen Uto, Finland. October
Galapagos flightless cormorant (Phalacrocorax harrisi) drying stunted wings after fishing, Fernandina Island, Galapagos, South America
Mute swan (Cygnus olor) landing on frozen lake, Richmond Park, London, UK. February
Common tern (Sterna hirundo) catching a mayfly, Hjalstaviken, Uppland, Sweden. Nordic Nature Photo Competition 2021 Highly Commended - Nordic Bird Photo of the Year category
A female kingfisher (Alcedo atthis) fishing, flying out of an ice hole in winter. Leeds, Yorkshire, UK. January
Black-headed gull (chroicocephalus ridibundus) diving into water, hunting fish / shellfish at sunrise. Baltic Sea, Poland. Sequence 1 of 2
White stork (Ciconia ciconia) flying back to its nest in Oak (Quercus robur) tree to feed its three large chicks, Knepp Estate, Sussex, UK, June
Looking down on Fulmar (Fulmarus glacialis) soaring above choppy sea, Orkney, Scotland, UK, May
Fulmar (Fulmarus glacialis) in flight against sky, Orkney, Scotland, UK, May
RF - Red admiral butterfly (Vanessa atalanta) feeding on rotten apples. Cornwall, England, UK. October. (This image may be licensed either as rights managed or royalty free.)
Campbell albatross (Thalassarche impavida), flying off Campbell Island (seen in background), January
Marsh fritillary butterfly (Euphydryas aurinia) nectaring on a Meadow buttercup (Ranunculus acris) flower in a chalk grassland meadow, Wiltshire, UK, June
Northern harrier (Circus hudsonius), two juvenile females hunting. North Park, Colorado, USA. August
Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February
Golden Eagle adult (Aquila chrysaetos) silhouetted in tree after sunset, Kuusamo, Finland, January
RF - Andean condor (Vultur gryphus) flying over Torres del Paine Massif. Torres del Paine National Park, Chilean Patagonia, Chile. March
White capped albatross (Thalassarche steadi) in flight at sea. Auckland Islands (Subantarctic), New Zealand, February
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Magnificent frigatebirds (Fregata magnificens) in low flight ovehead. Espanola Island, Galapagos, Ecuador, December
Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January
Sandhill crane (Grus canadensis) landing at sunset. North America. Highly honoured in the Art in Nature category of the Natures Best Photography Windland Smith Rice International Awards Competition
RF - Camouflaged moth (Agathia codina) Kinabalu National Park, Sabah, Malaysian Borneo. September
Scarce tortoiseshell butterfly (Nymphalis xanthomelas) on pussy willow, Luhanka, Keski-Finland, April. / Central Finland, Finland, April
Moth (Eumelea sp) Danum Valley, Sabah, Borneo
Emperor Moth (Saturnia pavoniella) male, on tree bark, Orvieto, Umbria, Italy, April
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
Galapagos / Flightless Cormorant (Phalacrocorax / Nannopterum harrisi) spreading its wings to dry, silhouetted against the sea. Fernandina Island, Punta Espinosa, Galapagos, Ecuador, April
RF- Great grey owl (Strix nebulosa) in flight. Kuhmo, Finland
Robin (Erithacus rubecula) taking off from snow, Hampstead Heath, London, UK, January
Jamess flamingo (Phoenicoparrus jamesi) walking with wings outstreched Laguna Colorada / Reserva Eduardo Avaroa, Altiplano, Bolivia
Western clubtail (Gomphus pulchellus) female, Groot Schietveld, Wuustwezel, Belgium, May
Cattle Egret (Bubulcus ibis) in breeding plumage, balancing on palm leaf, India
RF- Herring gull (Larus argentatus) in low flight over water, Flatanger, Nord Trondelag, Norway. August
RF- White stork (Ciconia ciconia) in flight, La Serena, Extremadura, Spain. March
RF- White-tailed Sea Eagle (Haliaeetus albicilla) turning in flight, Flatanger, Norway, August
Stellers Eagle (Haliaeteus pelagicus) in flight over snow, Hokkaido, Japan, February
Northern Chequered Skipper, (Carterocephalus silvicola), female on flower
Map Butterfly (Araschnia levana) feeding on anthemis, second generation, southern Finland, July
White naped cranes (Grus vipio) in flight, Kyushu, Japan
Common / Eurasian crane (Grus grus) displaying. Lake Hornborga, Sweden, April
