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Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
Wandering albatross {Diomedea exulans} flying over open ocean, South Atlantic
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016
Edwards Atlas Moth (Attacus edwardsii) in defensive posture, Bhutan, June
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Buzzard (Buteo buteo) in flight, portrait, Marlborough Downs, Wiltshire, UK. December
Great grey owl (Strix nebulosa) hunting over snow, woodland in background. Kuusamo, Finland. February
Golden eagle (Aquila chrysaetos) Sodankyla, Lokka, Finland, April
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Peregrine falcon (Falco peregrinus), adult male in flight. Bristol, UK. March
Lilac-breasted roller (Coracias caudata) stretching wings on acacia branch, Masai Mara Game Reserve, Kenya. November
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
Long eared owl (Asio otus) in flight, taking off from oak tree snag. Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Steppe eagle (Aquila nipalensis) in flight, Oman, November
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
Blood-vein moth (Timandra griseata) on fallen leaves, Nottingham, UK, August
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Barn Owl (Tyto alba) hunting / hovering, Somerset, UK, trained birdBarn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird
Siberian Jay (Perisoreus infaustus) alighting on a branch. Hamra, Sweden, September
Yellowhammers (Emberiza citrinella) coming in to land of a sheaf of oats in winter. Perthshire, Scotland, February
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Japanese / Red-crowned crane (Grus japonicus) three in flight, Hokkaido Japan February
Swallowtail butterfly (Papilio machaon), wings open, resting on plant stem. The Netherlands. August
A female kingfisher (Alcedo atthis) fishing through an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Kingfisher (Alcedo atthis) diving into water to catch fish, Near Bourne, Lincolnshire, England, UK. January
Bald eagle (Haliaeetus leucocephalus) flying over water in falling snow. Kachemak Bay, Alaska, USA. February
Ghost bat (Macroderma gigas) in flight, Pine Creek, Northern Territory, Australia
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Goshawk (Accipiter gentilis) flying in snow, Mainua Kajaani, Finland. March
European bee-eater (Merops apiaster) two perched on branch, Lower Silesia, Poland
Ruby-throated hummingbird (Archilochus colubris) male landing on Virginia creeper (Parthenocissus quinquefolia). Hill Country, Texas, USA