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Wide angle view of Common Swallowtail Butterfly (Papilio machaon) resting on Viper sWide angle view of Common Swallowtail Butterfly (Papilio machaon) resting on Vipers Bugloss / Blueweed (Echium vulgare) in alpine meadow
Clouded yellow butterfly (Colias crocea) perched on Ragwort flower. West Sussex, UK, September
Monarch butterfly (Danaus plexippus) hibernating, Mariposa Monarca Special Biosphere Reserve, central Mexico, January
White bellied woodstar (Chaetocercus mulsant) three hovering around red flowers, Guango private reserve, Papallacta Valley, Andean Cloud Forest, East slope, Ecuador
Purple gallinule (Porphyrio martinicus) moving across waterlily covered surface, Everglades National Park, Florida, USA, March
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
African moon moth (Argema mimosae) digital composite sequence, showing wings expanding after emerging from cocoon, photographed against a white background, orginating from Africa. Captive
Heath fritillary (Melitaea athalia) resting on scabious flower, Nordtirol, Austrian Alps, Austria, August
Tropical Butterfly (Adepha) on a leaf with wings spread. Danum Valley, Sabah, Borneo, Malaysia
Ichneumon wasp (Compsocryptus sp. ?) in flight, Williamson county, Texas, USA, NovemberIchneumon wasp (Compsocryptus sp.?) in flight, Williamson county, Texas, USA, November
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Common Swallowtail butterfly (Papilio machaon) resting on Meadow Clary (Salvia pratensis) flowers, on a white background. Nordtirol, Tirol, Austrian Alps, Austria, 1700 metres altitude, July
Jackdaws (Corvus monedula) abstract view of flock in flight, Derbyshire, UK, December. Non-ex
Golden eagle (Aquila chrysaetos) about to land on snow, Oulanka NP, Finland, February
Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June
European robin (Erithacus rubecula) in flight, Surrey, England
Red kite (Milvus milvus) in flight, Gigrin Farm, Powys, Wales, February
Mute swan (Cygnus olor) skidding along the surface of the water as it lands, Derbyshire, UK
Canada goose (Branta canadensis) landing at sunset, Derbyshire, UK
Madagascar fruit bat / flying fox (Pteropus rufus) Berenty Reserve, Madagascar (Digital composite)
Arctic tern (Sterna paradisaea) adult aggressively defending its nest from a perceived threat, Farne Islands, UK
Underside view of Manta Ray {Manta birostris} Mexico
Grey heron {Ardea cinerea} Al Ansab, Oman
Group of Rockhopper penguins {Eudyptes chrysocome} walking on beach, wings spread, Falkland Islands