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Juvenile Andean mouse opossum (Marmosops impavidus) eating a moth, El Oro, Ecuador
Great Green Macaw (Ara ambiguus) close up of feathers, El Manantial Macaw Sanctuary, Costa Rica. Captive
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April
Black Grouse (Tetrao tetrix) displaying with breath vapor at dawn, Utajarvi, Finland, April
Herring Gull (Larus argentatus) close up of juvenile in flight, Flatanger, Norway, October
Gentoo Penguin (Pygoscelis papua) Wiencke Island, Antarctic Peninsula, Antarctica
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Great grey owl (Strix nebulosa) taking off from snow, Kuhmo, Finland, February
Japanese / Red-crowned crane (Grus japonicus) three in flight, Hokkaido Japan February
Swallowtail butterfly (Papilio machaon), wings open, resting on plant stem. The Netherlands. August
A female kingfisher (Alcedo atthis) fishing through an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
RF - Ruppell's griffon vulture (Gyps rueppellii) drying its wings after a rain storm, perched on strangely gnarled tree (known locally as the Devil Tree')
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Kingfisher (Alcedo atthis) diving into water to catch fish, Near Bourne, Lincolnshire, England, UK. January
Bald eagle (Haliaeetus leucocephalus) flying over water in falling snow. Kachemak Bay, Alaska, USA. February
Whooper swans (Cygnus cygnus) pair engaged in courtship as snow falls, Myvatn lake, Iceland, March
RF - Atlantic puffin (Fratercula arctica) among sea campion (Silene maritima), Skomer Island, Pembrokeshire, UK. May
Close up of Malachite butterfly (Siproeta stelenes) wing detail. Captive
Great grey owl (Srix nebulosa) in flight about to land in snow, winter, Kuhmo, Finland, February
Greater flamingo (Phoenicopterus roseus) flock standing in lake as others fly overhead and one bird flaps its wings, Camargue, France, January
Blue and yellow macaw (Ara ararauna) in flight, Pantanal, Mato Grosso do Sul, Brazil
Small pearl-bordered fritillary (Boloria selene) roosting on ribwort plantain (Plantago lanceolata), Marsland mouth, Devonshire, UK. May
Wall brown butterfly {Lasiommata megera} basking on lichen covered stone wall, The Rumps, Polzeath, Cornwall, UK. April
Small tortoiseshell butterfly (Aglais urticae) basking on Cornish slate wall, Crantock, Cornwall, UK. June
Marbled white butterfly (Melanargia galathea) resting on common spotted orchid (Dactylorhiza fuchsii), Dunsdon, Devon, UK. July
Scarce chaser dragonfly (Libellula fulva) covered in dew, roosting on common sorrel (Rumex acetosa), Lower Tamar Lakes, Cornwall, UK. May
Osprey (Pandion haliaetus) fledgling strengthening wing in nest. Rasa Island Special Biosphere Reserve, Sea of Cortez, Mexico. April
Peacock katydid (Pterochroza ocellata) in defensive posture, Yasuni National Park, Orellana, Ecuador
Giant hummingbird (Patagona gigas) in flight, portrait, Cotopaxi National Park, Cotopaxi, Ecuador
Little egret (Egretta garzetta) landing on water, Ubrique, Southern Spain. June
Long-tailed tit (Aegithalos caudatus) feeding on icicle formed from sap of Painted maple tree (Acer pictum). Hokkaido, Japan. February
Atlantic puffin (Fratercula arctica) standing on rock with fish in beak, with wings spread, Machias Seal Island, Maine, USA. July
Great crested grebe (Podiceps cristatus) shaking chick from its back on lake, Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Hairy dragonfly (Brachytron pratense) male, resting on plant stem, Brackagh Moss National Nature Reserve, County Armagh, Northern Ireland, UK. June
Marsh fritillary butterfly (Euphydryas aurinia), imago stage, hanging from chrysalis, drying its wings. Focus Stacked. Captive
Close up of Madagascan silk moth (Antherina suraka) eyespot on wing showing hair-like scales. Focus Stacked. Captive
Abstract close-up of butterfly wing scales
Great white egret / Common egret (Ardea alba) preening feathers in shallow water of pond, Baie de Somme Nature Reserve, Hauts-de-France, France. September
Golden-edged owl-butterfly (Caligo uranus), butterfly captured for the collectors market, wing detail. Montes Azules Biosphere Reserve, Chiapas state, Mexico
Red grouse (Lagopus lagopus) in flight over moorland, Yorkshire, UK. September
Vermilion flycatcher (Pyrocephalus obscurus) male, courtship display, Texas, USA. May
Snowy owl (Bubo scandiaca) female, in flight in winter, Canada. January
White-tailed eagle (Haliaeetus albicilla) taking off from iceberg at sunrise, Shiretoko National Park, Hokkaido, Japan
Wandering albatross (Diomedea exulans) male, standing with wings spread in courtship display, Prion Island, South Georgia
Magnificent riflebird (Ptiloris magnificus alberti) male, courtship display, Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
