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Buzzard (Buteo buteo) in flight, portrait, Marlborough Downs, Wiltshire, UK. December
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
White-tailed eagle (Haliaeetus albicilla) with backlit wings, and water spray, Norway, October
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Calleta silkmoth (Eupackardia calleta), Texas USA Controlled conditions
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Great grey owl (Strix nebulosa) hunting over snow, woodland in background. Kuusamo, Finland. February
Osprey (Pandion haliaetus) carrying fish above river in Taipei, Taiwan
Golden eagle (Aquila chrysaetos) Sodankyla, Lokka, Finland, April
Saturniid moth (Leucanella hosmera), Chiriqui Province, Panama, South America
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Grey-headed flying-fox (Pteropus poliocephalus) spotting and flaring wings to come in to land on a branch. Yarra Bend Park, Kew, Victoria, Australia. ÔÇ¿November 2019
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch with one wing spread, Yarra Bend Park, Kew, Victoria, Australia.ÔÇ¿
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July
Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January
Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Indian Runner Duck, flapping wings
Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Great white egret (Ardea alba) group of three in winter, Kiskunsag National Park, Hungary. Winner of the Bird Category of Nature Photographer of the Year Awards 2018
Chinese moon moth (Actias dubernardi, female at rest amongst leaves. Dayaoshan, Jinxin, Guangxi, China
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Oleander Hawkmoth (Daphnis nerii) resting, Southern Sicily, Italy. March
Snowy owl (Bubo scandiacus) in flight, Ontario, Canada, January
Wild Buffons macaw x Scarlet macaw (Ara ambiguus x Ara macao) hybrid, Costa Rica
Fulvous forest skimmer, or Russet percher dragonfly (Neurothemis fulvia) female, Sai Kung, Hong Kong, China
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Straw-coloured fruit bat (Eidolon helvum), female flying carrying pup on front. Lamin, Gambia
