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Field of Giant plumrose anemones (Metridium giganteum) with a Snakelock anemone / Crimson sea anemone (Cribrinopsis fernaldi) in foreground, Alaska, USA, Gulf of Alaska. Pacific ocean
Sea otter (Enhydra lutris) floating on its back at surface among kelp, Alaska, USA. Gulf of Alaska, Pacific ocean
Wild Mustang horses running, Sand Wash Basin Herd Area, Colorado, USA
Wild Mustang horses fighting McCullough Peaks Herd Area, Wyoming, USA
Chestnut Mustang running in snow, at ranch, Shell, Wyoming, USA. February
Great Basin Bristlecone Pine (Pinus longaeva) fallen ancient tree, White Mountains, California, USA, March
Pika (Ochotona princeps) in scree rock pile, Sheepeaters Cliff, Yellowstone National Park, Wyoming, USA, September
American Red fox (Vulpes vulpes fulva) baby leaping on its disinterested mother. Grand Teton National Park, Wyoming, USA, May
Female Pintail (Anas acuta) with chicks, Katmai National Park and Preserve, Alaska, USA, July
American Red Fox (Vulpes vulpes) cubs playing with its mother. Grand Teton National Park, USA, June
American red squirrel (Tamiasciurus hudsonicus) collecting, Douglas Fir (Pseudotsuga menziesii) cones, Montana, USA
Yellow warbler (Dendroica petechia) collecting nesting material from Bulrush cattail (Typha sp) Bozeman, Montana, USA, June
Red winged blackbird (Agelaius phoeniceus) male singing, Bozeman, Montana, USA. April
American bison (Bison bison) walking through snow using tracks made by other animals, Yellowstone National Park, Wyoming, USA, February
Arctic ground squirrel (Spermophilus parryii) portrait, Denali National Park, Alaska, USA, September
Spruce grouse (Falcipennis canadensis) in forest, Denali National Park, Alaska, USA, September
Pika (Ochotona princeps) resting, Denali National Park, Alaska, USA, September
Bald eagle (Haliaeetus leucocephalus) in flight, Alaska, USA, February
Grizzly Bears (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
Grizzly Bear (Ursus arctos) asleep, Lake Clarke National Park, Alaska, September
Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Coastal brown bear (Ursus arctos) female in a meadow, cubs nearby, Lake Clarke National Park, Alaska, USA, July
Polar bear (Ursus maritimus) curious young male on the newly frozen pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Snow covered fishing boats in Sitka Harbour, Alaska. Photograph taken on location for BBC TV series Natures Great Events, March 2008
Spotted Owl (Strix occidentalis). Willamette National Forest, Oregon. June
California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Stone canyon jasper formed with cryptocrystalline quartz, California, USA
Polar bear (Ursus maritimus) silhouetted at waters edge at sunset, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, September. Vulnerable species
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Northern lights (Aurora borealis) glowing brightly over trees along Steese Highway, Cleary Point, outside Fairbanks, Alaska
Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
Tail fluke of a diving Humpback whale (Megaptera novaeangliae) misty coast in background, Alaska, USA. Gulf of Alaska, Pacific ocean
Three Sea otters (Enhydra lutris) resting on ice, Alaska, June
Grizzly Bear (Ursus arctos) eating a fish, Lake Clarke National Park, Alaska, August
Grizzly bear (Ursus arctos) female and nuzzling cub, Lake Clark National Park, Silver Salmon, Alaska. July
Sea otter (Enhydra lutris) female and pup playing. Alaska, USA, February
Panoramic of Point Lobos State Natural Reserve, Big Sur Coast, California, USA, September 2014
Bald eagle (Haliaeetus leucocephalus) flying over water in falling snow. Kachemak Bay, Alaska, USA. February
Rainclouds over Madison Valley, Beaverhead National Forest, Greater Yellowstone Ecosystem, Madision Mountains, Montana USA, July
Antelope Canyon, a slot canyon formed of Navajo sandstone and shaped by water erosion, Arizona, USA
Grey wolf (Canis lupus) pack resting on a snow ledge, Hayden Valley, Yellowstone National Park, USA. January
Red fox (Vulpes vulpes) resting in snow, Silver Gate, Yellowstone National Park, USA. January
American pine marten (Martes americana) standing on hind legs in deep snow, Yellowstone National Park, USA. January
Collie standing in snowy, frozen marsh, Eagle River, Alaska, USA. April