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Sea otter (Enhydra lutris) floating on its back at surface among kelp, Alaska, USA. Gulf of Alaska, Pacific ocean
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Golf-ball sized Painted frogfish (Antennarius pictus) waiting to ambush prey disguised as an orange sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Four Cedar waxwings (Bombycilla cedrorum) feeding on Crabapple (Malus sp) fruit, Ithaca, New York, USA, February
Adult Helmet Vanga (Euryceros prevostii) incubating a clutch of 3 eggs. Nest constructed in the fork of a Birds Nest Fern (Asplenium sp.) Masoala National Park, north east Madagascar
Great Blue shark (Prionace glauca) viewed from just below the surface, Pico Island, Azores, Portugal, Atlantic Ocean
European Brown Bear (Ursus arctos arctos), a large male taking a rest in a cloud of mosquitoes. Kajaani, Finland. June
Himalayan marmot (Marmota himalayana) sun bathing on rock, Mount Kailash, Tibet
Freshwater Dorado (Salminus brasiliensis) Recanto Ecologico, Rio da Prata, Bonito, Mato Grosso do Sul, Brazil
Japanese Macaque (Macaca fuscata) adult in the hot springs of Jigokudani, in the snow, Japan. February
Aggressive Mute swan (Cygnus olor) Upper Bavaria, Germany, Europe
Large white butterfly (Pieris brassicae), Vallee de l Eure (Eure Valley), Eure-et-Loir, France, August
Predatory bush cricket (Saga pedo) waiting for prey on Eryngium, Italy. June. Vulnerable species
Sand lizard (Lacerta agilis) male basking. Dorset, England, UK, May
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Blue sharks (Prionace glauca) cruise beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Red Admiral Butterfly (Vanessa atalanta) on Michaelmas daisy flowers. Dorset, UK, October 2012
RF - Marbled White butterfly (Melanargia galathea) resting on common spotted orchid, Dunsdon Nature Reserve, Devon, UK. July
RF- Pool Frog (Rana lessonae) sitting on white lily pad, Danube delta rewilding area, Romania
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
RF- Tiger (Panthera tigris) 14-month Lakshmi cub resting in tree, Bandhavgarh National Park, India. Endangered species
Grizzly bear (Ursus arctos) watching for Salmon at top of waterfall, Brooks Falls, Katmai National Park, Alaska, USA. July
Five Grizzly bears (Ursus arctos) standing at top of waterfall fishing for Salmon, Brooks Falls, Katmai National Park, Alaska, USA. July
Grizzly bears (Ursus arctos) on a waterfall, waiting for leaping Salmon, Brooks Falls, Katmai National Park, Alaska, USA. July
Two Pallas's cat (Otocolobus manul) kittens waiting at their den for their mother to return from hunting, Mongolia, June, 2017. Filmed for BBC series Big Cats
Jumping spider (Evarcha arcuata), female, waiting in curl of Oak (Quercus sp) leaf. Buckinghamshire, England. UK. September. Focus Stacked
Jumping spider (Evarcha arcuata), male, in alert pose, ready to jump. Buckinghamshire, England. UK. August. Focus Stacked
Two Oceanic manta rays (Mobula birostris) waiting to be cleaned by Clarion angelfish (Holacanthus clarionensis), Revillagigedo Islands, Mexico, Pacific Ocean
Female Red fox (Vulpes vulpes) waiting to be fed at the back entrance of a house, Hungary. January
Female Goldenrod crab spider (Misumena vatia) camouflaged on Yellow flag iris (Iris pseudacorus) flower waiting for prey to land, Kenfig National Nature Reserve, Glamorgan, Wales, UK, June
Darwins small ground finch (Geospiza fuliginosa), inspecting Galapagos giant tortoise for ticks, Isabela Island, Galapagos, South America
Flower spider (Thomisus spectabilis) an ambush predator waiting on daisy flower for unsuspecting pollinators, Toowoomba, Queensland, Australia
RF- Morpho butterfly displaying on leaf (Morpho achilles). Amazonia, Ecuador
RF- Red Admiral butterfly (Vanessa atalanta) perched on Bigtooth Maple (Acer grandidentatum). Lost Maples State Park, Hill Country, Central Texas, USA. November
RF- Free range Domestic pig (Sus scrofa domesticus) waiting for feed, UK, August 2010
Head portrait of grey Andalusian stallion in dark barn. Northern France, Europe
Brown hare (Lepus europaeus) three adults in snow covered field during a snow fall, Derbyshire, UK, February
Leopard (Panthera pardus fusca), watching from rock, Rajasthan, India
RF - Strawberry Poison Frog (Oophaga pumilio). Central Caribbean foothills, Costa Rica
Bighorn Sheep (Ovis canadensis) on the hillside, ruminating, Yellowstone National Park, Wyoming, USA. January 2011
Bison (Bison bison) standing steam from geothermal springs, Yellowstone National Park, Wyoming, USA, February
RF- Very young Florida manatee (Trichechus manatus latirostrus) Manatees are often born with very wrinkled skin. Crystal River, Florida, USA. February 2010
RF- Red squirrel (Sciurus vulgaris) portrait, in snow, Cairngorms National Park, Scotland, March 2007
RF- Rainbow trout (Oncorhynchus mykiss) in lake, Capernwray, Lancashire, UK, July. Did you know? Rainbow trout were indigenous only to Alaska and east Asia
Willow grouse (Lagopus lagopus) Kiilopaa, Inari, Finland, January
Long-jawed orbweaver (Metellina segmentata) in web at sunset. Dunwich Heath, Suffolk, UK, September