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California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Northern cardinal (Cardinalis cardinalis) male perched amid snow-covered branches, New York, USA, February
Red-breasted sapsucker (Sphyrapicus ruber) adult feeding on tree sap from shallow wells it has drilled in bark of Curl-leaf Mountain-mahogany (Cercocarpus ledifoius), Lee Vining Canyon, California
Crested caracaras (Caracara cheriway), midair aggressive interaction between two for access to perch, Martin Refuge, Rio Grande Valley, Texas, USA, March
Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April
Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May
Tricolored heron (Egretta tricolor) adult in breeding plumage capturing fish by half-running half-flying across waters surface and stabbing with its bill, Viera Wetlands, Brevard County, Florida
Royal tern (Sterna maxima) perching on a sand castle on a beach, Fort De Soto Park, St. Petersburg, Florida, USA, March
Ruby-throated Hummingbird (Archilocus colubris), male flying in to feed from cardinal flowers (Lobelia cardinalis) New York, USA. August
Abstract interpretation of Sandhill Cranes (Grus canadensis) at roost during spring migration in March, Rowe Sanctuary, Kearney, Nebraska, USA. March. Long exposure
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Sea otter (Enhydra lutris) group of four resting among sea ice, Alaska, USA, June
Glowworm beetle (Phengodes sp. ) male flying, Tuscaloosa County, Alabama, USA Controlled conditionsGlowworm beetle (Phengodes sp.) male flying, Tuscaloosa County, Alabama, USA Controlled conditions. July
Humpback Whale (Megaptera novaeangliae) blowing or spouting and fluking, Southeast Alaska, USA, August
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Giant sequoia (Sequoiadendron giganteum) trees in Sequoia National Park, California, USA, September 2014
Young Polar bear (Ursus maritimus) standing on hing legs, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Green june beetle (Cotinis nitida) in flight Williamson County, Texas, USA Controlled conditions. October
Florida manatees (Trichechus manatus latirostris) gathering at Three Sisters Spring in Crystal River, Florida, USA
Moose (Alces alces) female with twin calves, Baxter State Park, Maine, USA, June
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
English springer spaniel in Rhododendron. Haddam, Middlesex, Connecticut, USA
Weimaraner in front of Rhododendron flowers, Haddam, Connecticut, USA. May
Bichon frise beside Colorado blue columbine (Aquilegia coerulea) flowers. Old Lyme, Connecticut, USA. May
Bichon frise standing on rock amongst Rhododendron flowers. Haddam, Connecticut, USA. May
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
Purebred Berkshire piglet amongst autumn foliage, Smithfield, Rhode Island, USA
Collared peccary (Tayassu tajacu), adult drinking, South Texas, USA. May
Polar bear (Ursus maritimus) swimming at surface of Beaufort Sea, Alaska, USA
Polar bear (Ursus maritimus) sow standing along snow covered Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Polar bear (Ursus maritimus) juvenile crossing over newly forming pack ice, during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
RF - Two Gypsy vanner geldings standing together looking away, Greyrock, Wyoming, USA. June
Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA
Fillys eye, wild Mustang horse at Black Hills Wild Horse Sanctuary, South Dakota, USA
Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August
Two Gypsy vanner geldings in wildflower pastures, Wyoming, USA. June 2014
Wild Mustang horse rearing, Pryor mountains, Montana, USA. June
RF - Head portrait of grey Andalusian mare with long mane in snow, Berthoud, Colorado, USA. January
RF - Wild pinto Mustang stallion running in snow, Black Hills Wild Horse Sanctuary, South Dakota, USA. January
Black bears (Ursus americanus) standing on back legs, fighting, Minnesota, USA, June
