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Leather Coral (Sinularia flexibilis). Komodo National Park, Indonesia
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Magnificent sea anemone (Heteractis magnifica) with Anemone fish within tentacles, Indonesia. Indo-Pacific
Broadclub cuttlefish hunting technique - mesmorises fish by pulsating colour along body
Crown of thorns starfish {Acanthaster planci} Andaman Sea, Thailand
Grey whale (Eschrichtius robustus) eye, Magdalena Bay, Baja California, Mexico, February. Winner in the Ocean Views category of the 2018 Natures Best Winland Smith Rice Awards
Indo-Pacific humpback dolphin (Sousa chinensis) with fishing net damage around throat, Pearl River Estuary, Lantau Island, Hong Kong, China
RF - Soft coral (Dendronephthya sp. ) growing on sea bed. West Papua, IndonesiaRF - Soft coral (Dendronephthya sp.) growing on sea bed. West Papua, Indonesia
Paddle-flap scorpionfish (Rhinopias eschmeyeri) Puerto Galera, Philippines
Necklace seastar (Fromia monilis) on Sea cucumber (Bohadaschia argus) Yap, Micronesia
Broadclub cuttlefish (Sepia latimanus) in translucent egg casing on the reef, Philippines, Asia
Three spot damselfish (Dascyllus trimaculatus) with Sea anemone home, Yap, Micronesia
Common anemonefish (Amphiprion perideraion) with eggs in Magnificent sea anemone (Heteractis magnifica) Yap, Micronesia
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha ab Mahmud, Sinai, Egypt. Red Sea
Southern stingray (Dasyatis americana) large female foraging over seagrass in shallow water. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Crowned dwarf caiman (Paleosuchus trigonatus) male resting on a log. Yasuni National Park, Orellana, Ecuador
Flower tree soft corals (Umbellulifera sp. ), Triton Bay, near Kaimana, West PapuaFlower tree soft corals (Umbellulifera sp.), Triton Bay, near Kaimana, West Papua, Indonesia
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluespine unicornfish (Naso unicornis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Veined octopus (Amphioctopus marginatus) between clam shell halves. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Blue whale (Balaenoptera musculus) diving beneath the surface. Indian Ocean, Sri Lanka
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean
Brydes whale (Balaenoptera edeni) Trincomalee, Eastern Province, Sri Lanka, Bay of Bengal, Indian Ocean
Humboldt squid (Dosidicus gigas) cannibalising another squid of the same species at night off Loreto, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Green Sea Turtle (Chelonia mydas) returning to sea after Egg laying, Raine Island, Australia. Raine Island is the largest and most important green sea turtle nesting area in the world, with over 14
Rainbow lorikeet (Trichoglossus haematodus) Queensland, Australia
Cairns birdwing butterfly (Ornithoptera priamus) male on fern leaf and Blue Mountain Swallowtail butterfly in flight, Atherton Tablelands, Queensland, Australia, October
A male Leafy Seadragon (Phycodurus eques) carries a new batch of eggs on his tail. Wool Bay Jetty, Edithburgh, Yorke Peninsula, South Australia, November
Dwarf Minke whale (Balaenoptera acutorostrata) Great Barrier Reef, Australia. Coral Sea
Florida manatee (Trichechus manatus latirostrus), Three Sisters Springs, Crystal River, Florida, United States
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Raggiana bird of paradise (Paradisaea raggiana) male displaying to female, Papua New Guinea, Varirata NP
Snorkeller with shoal of Piraputanga (Brycon hilarii), Rio Olha d Agua at Cabeceira do Prata, Bonito, Mato Grosso do Sul, Brazil
Ceiba / Kapok tree (Ceiba trichistandra) low angle view, Macara, Loja, Ecuador
Sumatran pit viper (Trimeresurus sumatranus) female in rain forest understorey. Danum Valley, Sabah, Borneo
White spotted eagle ray (Aetobatus narinari) feeding by digging in the sand and seagrass for invertebrates. East End, Grand Cayman, Cayman Islands, Caribbean Sea
Portrait of Blacktip grouper (Epinephelus fasciatus) resting in Barrel sponge (Xestospongia testudinaria) Mabul Island, Sabah, Borneo, Malaysia. Sulawesi Sea
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Queen conch (Strombus gigas) extending its proboscis and stalked eyes at it moves across the sand by coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Large male Napoleon wrasse (Cheilinus undulatus) yawning. Ras Mohammed, Sinai, Egypt. Red Sea
School of Longnose parrotfish (Hipposcarus harid) swarming over reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Southern stingray (Hypanus americanus) female swimming over a shallow sand bank, under morning sun, North Sound, Grand Cayman, Cayman Islands. British West Indies, Caribbean Sea
Shallow coral reef flat with corals (Pocillopora damicornis, Acropora humilis and Pocillopora verrucosa) in evening light. Gordon Reef, Strait of Tiran, Sinai, Egypt. Gulf of Aqaba, Red Sea
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea