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Scots Pines (Pinus sylvestris) and flowering heather moorland at sunrise, Abernethy, Cairngorms National Park, August 2016
Old trees in the Syndicate Forest National Park, Dominica, Eastern Caribbean. February 2017
Red shanked douc langur (Pygathrix nemaeus) Son Tra Nature Reserve, Vietnam
Colombian red howler monkey (Alouatta seniculus) mother and baby in tree. Northern Colombia
Hawthorn tree and granite outcrop at Holwell tor, sunrise and mist, Dartmoor National Park, Devon, UK. October 2018
Pileated Woodpecker (Dryocopus pileatus) male, at feeding excavation in tree trunk in winter, Freeville, New York, USA, January
Common redpoll (Carduelis flammea), male perched in alder (Alnus rugosa) during a snowstorm in winter, New York, USA, January
Stream in spate in native oak woodland in summer, Clonaig, Kintyre, Argyll, Scotland, UK, July 2015
Montane rainforest, Hienghene, New Caledonia
Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Freshwater pools on Waterfall Creek, at the top of Gunlom Falls. Mary River, Kakadu National Park, Northern Territory, Australia. March 2014
Lion (Panthera leo) family climbing and resting in tree. Serengeti National Park, Serengeti, Tanzania
Lion (Panthera leo), female in fork of tree. Serengeti National Park, Serengeti, Tanzania
Beech woodland near Blackwater Brook. New Forest National Park. November 2014
Peach trees (Prunus persica) in flower, Fruiturisme Tourism Experience, Lleida, Catalonia, Spain. March 2018
Lemon shark pups (Negaprion brevirostris) spend the first 5-8 years of their life in mangrove forests. The tangle of roots provides protection from predators
Composite image of Fireflies synchronize display just after first shower of monsoon, on moonless night. India
Ivy (Hedera helix) covered tree trunks in Knight Wood, Hampshire, UK, October
Giant honey bee nest (Apis dorsata) up in a giant Mengaris tree (Koompassia excelsa) dipterocarp forest in Danum Valley, Sabah, Borneo, Malaysia
Long-tailed tit (Aegithalos caudatus caudatus) placing feather in nest, Finland. April
Blackcap (Sylvia atricapilla) female and two males huddling together for warmth before going to sleep, Finland, May
Lesser spotted woodpecker (Dendrocopos minor), male at nest hole, Finland, June
Willow (Salix) plantation, Cuenca, Castilla-La Mancha, Spain, December
Misty morning view of Exmoor countryside from Winsford Hill, Exmoor National Park, Somerset, UK. October 2016
Rock formations at Brimham Rocks created by variable erosion of soft and hard layers of carboniferous age millstone grit, Harrogate, Yorkshire, England, UK, October 2016
Strangler fig (Ficus sp) growing over Hoop Pine tree(Araucaria cunninghamii) with branches knotting round each other, Morans Falls Track, Green Mountains
Banyan tree (Ficus macrophylla) Valley of the Shadows, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Birch tree (Betula) growing in Llyn Padarn, Llanberis, Snowdonia National Park, North Wales, UK, March
Hazel Grouse (Tetrastes bonasia) in snow covered trees, Suomussalmi, Finland, January
Red panda (Ailurus fulgens) family, adult female and two subadult juveniles, rest together in the canopy of the cloud forest. Singalila National Park, West Bengal, India
Autumn colours and snow covered mountains Baima Snow Mountain Nature reserve, Yunnan, China, October 2017
Chinese Lanterns (Physalis alkekengi), Hydrangea flowers, Poplar leaves (Populus) and Honesty skeletons (Lunaria annua), silhouettes on white background
Newfoundland dog, male standing in snow. Killingworth, Connecticut, USA
Aerial view of lowland Amazonia rainforest in morning mist. Manu Biosphere Reserve, Amazonia, Peru. November 2017
Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017
Red ruffed lemur (Varecia rubra) female in understorey of lowland rainforest. Masoala National Park, north-east Madagascar
Silky sifaka (Propithecus candidus), female with baby sitting in rainforest understorey. Mid-altitude montane rainforest, Marojejy National Park, north-east Madagascar
Waterfall in Fjaorargljufur canyon, Kirkjubaejarklaustur, Iceland, August 2017
Riisitunturi in winter with Aurora Borealis, Kuusamo, Lapland, Finland. January 2016
Jasna Lakes in autumn, Kranjska Gora, Julian Alps, Slovenia. October 2014
Church of St. Primoz, Jamnik, Slovenia. July 2017
Morel mushroom (Morchella sp) Sierra de Grazalema Natural Park, southern Spain, May
Red deer (Cervus elaphus) stag bellowing, Los Alcornocales Natural Park, southern Spain, September
River and beech woodland (Fagus sylvatica) Muniellos National Park, Asturias, Spain. October
Holm oaks (Quercus ilex) two on hillside, Arcos de la Frontera, southern Spain. March
European white birch (Betula pubescens) woodland in winter, Spain, March 2016
Sessile oaks (Quercus petraea) and Beech trees (Fagus sylvatica) in winter, Muniellos National Park, Asturias, Spain. April
Tawny owl (Strix aluco) sleeping, Cadiz, Andalusia, Spain, February