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Western Grebe (Aechmophorus occidentalis) swimming, Guerrero Negro, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Salmon (Salmonidae) fry with egg case still attached, Lake Kuril, Kamchatka, Far East Russia, January
Bewicks swan (Cygnus columbianus) in flight with rainbow, Gloucestershire, England, UK, February
RF - Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean
Brown hare (Lepus europaeus) three adults in snow covered field during a snow fall, Derbyshire, UK, February
Fulmars (Fulmarus glacialis) on water, feeding at a fish processing plant outfall. Grundafjordur, Iceland, March
Greater flamingos (Phoenicopterus ruber) in flight over sand dune, Donana National Park, Andalusia, Spain, March
Four Yellow-billed storks (Mycteria ibis) standing in water, Lake Nakuru, Nakuru National Park, Kenya
Female humpback whale (Megaptera novaeangliae) with its very young calf at the surface, Rurutu island, Australs Archipelago, French Polynesia. Pacific Ocean
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
European Herring Gull (Larus argentatus) flock in flight at sunset Norway, May
Grey herons (Ardea cinerea) standing in water a dawn, Puszaszer, Hungary, July
Persian kittens among snowy flowers
Three Silver studded blue butterflies (Plebejus argus) covered in dew, Klein Schietveld, Brasschaat, Belgium
Lion (Panthera leo) cubs playing, Masai Mara Game Reserve, Kenya
Wild Mustang pinto foal nuzzling up to mother, Sand Wash Basin Herd Area, Colorado, USA
White-tailed deer (Odocoileus virginianus) in snow, Acadia National Park, Maine, USA, February
Herd of Impala (Aepyceros melampus) silhouetted at sunset, Ngorongoro Conservation Area, Tanzania
Yellow Mongooses (Cynictis penicillata) standing alert, Kgalagadi National Park, South Africa
Komodo dragons (Varanus komodoensis) in forest clearing, Komodo Island, Indonesia
Cheetah (Acinonyx jubatus) mother and juveniles, Masai-Mara Game Reserve, Kenya. Vulnerable species
Atlantic puffins (Fratercula arctica) at dusk, Fair Isle, Shetland, July. Bookplate from Danny Greens The Long Journey North
Mature Bighorn Rams (Ovis canadensis) on high mountain pass. Jasper National Park, Alberta, Canada, October
Sequence of a breaching Southern right whale (Eubalaena australis) Golfo Nuevo, Peninsula Valdes, UNESCO Natural World Heritage Site, Chubut, Patagonia, Argentina, Atlantic Ocean
Puffins (Fratercula arctica) among coastal thrift. Saltee Islands, Ireland, UK, July
Bengal Tiger (Panthera tigris tigris) mother and cub, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle
Bengal Tigers (Panthera tigris tigris) resting in the shade national park, India, taken on location for Tiger - Spy in the Jungle March 2007
Domestic dog, St. Bernard / Alpine Mastiff, three on snow in Alps, France
Herd of Camargue bulls, Camargue, France, May
A group of Pure Arab, Shagya Arab and East Bulgarian fillies running in snow, Kabiuk National Stud, Shumen, Bulgaria
Lesser flamingos (Phopeniconaias minor) flock, aerial view, Natron lake, Rift valley, Tanzania, Africa
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
Polar bear (Ursus maritimus) mother and large second year cub swim across calm water, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
A band of semi-feral Quarter mares and foals running across a creek, Estancia Don Amerigo, Chaco, Paraguay, January 2012
Asiatic lionesses and cubs (Panthera leo persica) drinking from pool, Gir Forest NP, Gujarat, India. Water is scarce in the Gir
Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May
Giraffe (Giraffa camelopardalis) three standing together, silhouetted at dusk, Okavango Delta, Botswana
Flock of Cape petrels (Daption capense capense) in flight against the sea. Against the sun. South Shetland Islands, Antarctica, January
Common dolphins ( Delphinus delphis) swimming below the surface, Atlantic Ocean, Portugal, October
A pod of Bottlenose dolphins (Tursiops truncatus) swimming into the Sado river, Portugal, July
House sparrows (Passer domesticus) perched on building, Morocco, March
Hand reared Whooper Swans (Cygnus cygnus) in flight. Birds imprinted and trained by Lloyd and Rose Buck for filming purposes. Somerset, UK, January 2011
Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October
Alpine Chough / Yellow-billed Chough (Pyrrhocorax graculus) flock in flight against mountains, Dolomites, Italy. Digital composite
Roe Deer (Capreolus capreolus) swimming across River Allier, France, February
Black-legged Kittiwake (Rissa tridactyla) on ice. Floe edge, Arctic Bay, Nunavut, Canada, April