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Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;
Two Black woodpeckers (Dryocopus martius) males on tree trunk in territorial dispute, Peerdsbos, Braschaat, Belgium. April
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Asian black bear (Ursus thibetanus) scent marking tree by rubbing back on it, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it, Land of the Leopard National Park, Russian Far East. Endangered
Lion (Panthera leo) male, scent marking its territory at dawn, Ngorongoro Conservation Area / Serengeti National Park boundary, northern Tanzania
Northern parula (Parula americana) male singing from flowering crabapple (Malus sp)Northern parula (Parula americana) male singing from flowering crabapple (Malus sp.) tree in spring, near Salamanca, New York, USA, May 2022
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Male Yellowface pike-blenny (Chaenopsis limbaughi), living in abandoned tubeworm holes in sand, flaring its dorsal fin in threat display, Dominica, Eastern Caribbean
Dippers (Cinclus cinclus) standing on ice, ready to fight over feeding territory, Kuusamo, Finland, February
Dippers (Cinclus cinclus) fighting over feeding territory, Kuusamo, Finland, February
Two male Darjeeling bush frogs (Raorchestes Annandalii) fighting over territory, Kalimpong, West Bengal, India
Golden damsel (Amblyglyphidodon aureus) male aggressively defends his territory on a coral reef. Piaynemo Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Mute swan (Cygnus olor) in a threathening territorial pose and attacking a rival Lake Geneva, near Geneva, Switzerland, March
Iberian lynx (Lynx pardinus) sniffing scent on dead branch, Parque Natural Sierra de Andujar, Andalucia, Spain. January
Bengal Tiger (Panthera tigris) Arrowhead patrolling his territory, Ranthambore National Park, India
Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA