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Aerial view of nesting Gannets (Morus bassanus) and swirling seas, Hermaness, Shetland, Scotland, UK, October
Darter ( Anhinga melanogaster) drying wings, Keoladeo NP, Bharatpur, India
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Shag (Phalacrocorax aristotelis) two heads showing above rock, Vardo, Norway March
Northern gannet (Morus bassanus), Feeding frenzy, Noss NNR, Shetland, Scotland, UK
Great cormorant (Phalacrocorax carbo) in snow. Netherlands. January
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
RF - Gannet (Morus bassanus) close-up portrait with nesting material in beak, , Saltee Islands, Ireland. June
Gannet (Morus bassanus) adult turning in the air as it begins to plunge dive. Shetland Islands, Scotland, UK, July
Great Frigatebird (Fregata minor) in flight over Midway Atoll. Midway
Gannet (Morus bassanus) adult wing stretching and looking up, Shetland Islands, Scotland, UK, September
Gannet (Sula bassana) in water, Shetland, UK. July
Northern gannets (Morus bassanus) diving for fish. Bempton, Yorkshire, UK. August
Brandts cormorant (Phalacrocorax penicillatus) swimming beneath an oil rig. Eureka Rig, Los Angeles, California, United States of America. North East Pacific Ocean
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
Booby (Sula nebouxii) looking at camera underwater, Galapagos Island
Red-footed booby (Sula sula) perched on Palo santo tree (Bursera graveolens), with flowering Yellow cordia (Cordia lutea) in background. Galapagos Islands, Ecuador
Northern gannets (Morus bassanus) breeding colony nesting on sea cliffs, Hermaness, Shetland, Scotland, UK. July
Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp)Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp.) being followed by a California sea lion (Zalophus californianus), Espiritu Santo Island
Magnificent frigatebirds (Fregata magnificens) pair in courtship display, male with gular sac inflated, North Seymour Island, Galapagos National Park, Galapagos Islands
Red-footed booby (Sula sula) chick and parent perched on branch near nest, Genovesa Island, Galapagos Islands, Ecuador
Red-footed booby (Sula sula) perched on branch, Genovesa Island, Galapagos Islands, Ecuador
Great frigatebird (Fregata minor) attempting to steal food from Nazca booby (Sula granti) feeding its chick, Genovesa Island, Galapagos Islands, Ecuador
Male Great frigatebird (Fregata minor) in breeding condition with semi-inflated throat pouch, Christmas Island, Australia
Blue-footed booby (Sula nebouxii), incubating eggs with webbed feet, Seymour Island, Galapagos, South America
Galapagos flightless cormorant (Phalacrocorax harrisi) drying stunted wings after fishing, Fernandina Island, Galapagos, South America
Adult Shags (Phalacrocorax aristotelis) with young, gathered on rocks, Treshnish Isles, Southern Hebrides, Scotland, UK, Atlantic Ocean. August
Cormorant (Phalacrocorax carbo) swimming on pond, Valkenhorst Nature Reserve, The Netherlands, August
Young and adult Shags (Phalacrocorax aristotelis) gathered on rocks in the sea. Treshnish Isles, Southern Hebrides, Scotland, UK. Sea of Hebrides, North East Atlantic Ocean
Northern gannets (Morus bassanus) pair displaying amongst campion, Bempton Cliffs, Yorkshire, UK, May
Cormorants (Phalacrocorax carbo) in early morning mist, Oisterwijkse Bossen en Vennen nature reserve, Netherlands, November
Flightless cormorant (Phalacrocorax harrisi), comoing ashore after fishing trip, Fernandina Island, Galapagos Islands
Blue-footed booby (Sula nebouxii) pair with chicks, blowhole in background, Punta Suarez, Espanola Island, Galapagos Islands
Brown booby (Sula leucogaster), Los Islotes, Espiritu Santo Archipelago National Park, Sea of Cortez (Gulf of California), Mexico, September
Blue-footed booby (Sula nebouxii), diving, Santa Cruz Island, Galapagos Islands
Bronze whaler shark (Carcharhinus brachyurus) group feeding alongside Cape gannet (Morus capensis) during sardine run. Wild Coast, Eastern Cape, South Africa. June
Split-level of Cormorants (Phalacrocorax sp. ) standing on rocks with Sea lettuce (Ulva spSplit-level of Cormorants (Phalacrocorax sp.) standing on rocks with Sea lettuce (Ulva sp.) seen underwater, Hussar Bay, Nigei Island, Queen Charlotte Strait, British Columbia, Canada. September
Red-footed booby (Sula sula) landing among salt bushes, looking for nesting materials, with Frigate birds behind, Genovesa Island, Galapagos