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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Scots pine (Pinus sylvestris) trees reflected in Loch Maree at sunrise with Slioch in background, Wester Ross, Scotland, UK, November 2014
English oak tree (Quercus robur) at sunset, Monmouthshire Wales UK, March
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Super moon rising above Glastonbury Tor with people watching, Somerset, England, UK, 27th September 2015
Knoydart and Loch Nevis from Sgurr Cos na Breachd-laoidh with sun shining through cloud, Lochaber, Scotland, UK, September 2016
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Silhouette of Common peafowl (Pavo cristatus) perched in a tree at dawn. Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Chinese lanterns (Physalis alkekengi) skeletons and Honesty seed pods (Lunaria annua)
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
Star trails above Scots pine (Pinus sylvestris) woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, March 2017
Scots pine (Pinus sylvestris) trees reflected in Loch Maree at dawn with Slioch in background, Wester Ross, Scotland, UK, November 2014
Two Rooks (Corvus frugilegus) silhouetted as they perch on a tree branch at their roost site at sunset, Gloucestershire, UK, February
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Fallow deer (Cervus dama) stags at sunset, Holkham Park, North Norfolk, UK, January
Sea stacks and cliffs at sunset at Eshaness / Esha Ness, peninsula in Northmavine on the island of Mainland, Shetland Islands, Scotland, UK, May 2018
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
Helicopter flying over volcanic eruption near Eyjafjallajoekull glacier, Iceland, 24th March 2010. Volcano previously dormant since 1821
Oak tree (Quercus robur) silhouetted against night sky with stars, Brecon Beacons National Park, Wales, UK. December
Crested newt (Triturus carnifex) Gelderland, The Netherlands, April. Winner of the Amphibians category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Red fox (Vulpes vulpes) silhouetted in mixed beech-Douglas fir forest. Freiburg, Black Forest National Park, Germany, October
Sun rays shining through clouds onto Loughrigg Fell and hills of the Lake District National Park, in evening. Ambleside, Cumbria, England, UK. April 2020
Scalloped hammerhead shark shoal (Sphyrna lewini) Wolf Island, Galapagos
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
Quiver trees (Aloidendron dichotomum) silhouetted against dawn sky, Namibia, May
Avielochan at sunrise, Cairngorms National Park, Scotland, UK, August
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Humpback Whale (Megaptera novaeangliae) tail fluke at surface as whale dives, in the late afternoon light, Andenes, Andoya island, North Atlantic Ocean, Norway Janurary
Teasel (Dipsacus fullonum), Hedge parsley (Torilis) and Allium seedhead silhouettes on white background
Short-eared owl (Asio flammeus) at sunset, Vendee, France, February
Polar bear (Ursus maritimus) silhouetted against setting sun, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Silhouetted Dartmoor pony (Equus caballus) at sunrise, Combestone Tor, Dartmoor National Park, Devon, England, UK, January
RF- Small group of Pink-footed geese (Anser brachyrhynchus) in flight at dawn, The Wash Estuary, Norfolk, England, UK, October 2011
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Topi (Damaliscus korrigum) at sunset, Masai-Mara Game Reserve, Kenya
Red deer stag (Cervus elaphus) silhouetted against sky at dusk, Cheshire, October 2014
Red deer (Cervus elaphus) stag bellowing during rut, at night with full moon, Derbyshire, UK
Gemsbok (Oryx gazella) silhouetted at dawn, Kalahari Desert, Botswana
Stag beetle (Lucanus cervus) male silhouetted against leaf, controlled conditions
English oak tree (Quercus robur) at night silhouetted against circle of star trails, Brecon Beacons Wales, UK. January 2017
Sunset over savanna landscape image with a lone (Acacia) tree, Masai Mara NR, Kenya