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Espanola lava lizard (Microlophus delanonis) female with Waved albatross (Phoebastria irrorata) behind. Punta Suarez, Espanola Island, Galapagos
Red-footed booby (Sula sula) landing among salt bushes, looking for nesting materials, with Frigate birds behind, Genovesa Island, Galapagos
Nazca booby (Sula granti) on coast, with Vampire ground finches (Geospiza septentrionalis). These finches peck at birds, feeding on blood when other food sources are scarce, Wolf sland, Galapagos
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
Nazca booby (Sula granti) on nest. Genovesa Island, Galapagos. February 2017
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) pair mutual preening in courtship. Genovesa Island, Galapagos Islands, Ecuador, June
Galapagos Petrel (Pterodroma phaeopygia) on debris from nesting hole. Critically endangered. Santa Cruz Highlands, Galapagos Islands, Ecuador, June
Waved Albatross (Phoebastria irrorata) and Blue-footed Booby (Sula nebouxii) squabbling over nesting sites. Galapagos Islands, Ecuador, June
Balearic shearwater (Puffinus mauretanicus) at night inside a breeding colony cave, Des Conills islet, Malgrats islands (ZEPA, Zona de Especial Proteccion para las Aves), Majorca
Atlantic puffin (Fratercula arctica) in breeding plumage flapping wings on sea cliff top in seabird colony, Scotland, UK, May. Digital composite
Northern fulmar (Fulmarus glacialis) in flight with waves, Arctic Sea. Commended in the Biophoto Competition 2019
Puffin (Fratercula arctica) landing in snow, Norway, March
White capped albatross (Thalassarche steadi) in flight at sea. Auckland Islands (Subantarctic), New Zealand, February
Black-browed albatross (Thalassarche melanophris) in flight over sea at sunrise. South Atlantic Ocean between The Falklands and South Georgia. November
Southern royal albatross (Diomedea epomophora) sub-adults, group displaying in tussock grassland. Campbell Island, New Zealand. February
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Magnificent frigatebirds (Fregata magnificens) in low flight ovehead. Espanola Island, Galapagos, Ecuador, December
Flightless Cormorant (Nannopterum / Phalacrocorax harrisi) with nesting material. Banks Bay, Isabela Island, Galapagos Islands, Ecuador, December
Blue-footed Booby (Sula nebouxii) on guano covered rocks. Santa Cruz Island, Galapagos, June
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Gannet (Morus bassanus) adults and a youngster silhouetted against an orange red sky as another adult prepares to land. Shetland Islands, Scotland, UK. September
Puffin (Fratercula arctica) silhouetted at sunset on Hermaness, Shetland, Scotland, UK, June
Gannet (Morus bassanus) in flight is silhouetted against the setting sun. Shetland Islands, Scotland, UK, August
Gannet (Morus bassanus) chick with fluffy with mouth open. Shetland Islands, Scotland, UK, August
Gannet (Morus bassanus) chick in a line of adults. Shetland Islands, Scotland, UK, August
Balearic shearwater (Puffinus mauretanicus) in flight over Irish sea off Pembrokeshire coast, Wales, UK, August 2010, endangered species
Northern gannet (Morus bassanus) colony in flight over Bass Rock at sunrise, Firth of Forth, Scotland, August 2009
Black Skimmer (Rynchops nigra), skimming with its open beak just below the water sBlack Skimmer (Rynchops nigra), skimming with its open beak just below the waters surface in search of prey, Bolsa Chica Ecological Reserve, California
Black-browed Albatross (Thalassarche melanophris) resting close up of eye. West Point Island, Falkland Islands, October
Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Red Footed Booby (Sula sula) couple, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Laysan Albatross (Phoebastria immutabilis) in nest, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Northern gannets (Morus bassanus). Bass Rock, Scotland, UK. August
Atlantic Puffin (Fratercula arctica) covered by mud from its partner digging out their nesting chamber / burrow, Skomer Island, Wales, UK, April
Atlantic Puffin (Fratercula arctica) swimming, split level view, Skomer Island, Wales, UK, July
Atlantic Puffins (Fratercula arctica) in flight in mist, Skomer Island, Wales, UK, August
Gannet (Morus bassanus) nearest out of focus, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Gulls flying above two Killer whales / Orcas (Orcinus orca) surfacing, Tysfjord, Norway, November
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
Galapagos / Flightless Cormorant (Phalacrocorax / Nannopterum harrisi) spreading its wings to dry, silhouetted against the sea. Fernandina Island, Punta Espinosa, Galapagos, Ecuador, April
Razorbill (Alca Torda) on cliff, Farne Islands, Northumberland, England, UK, June
Razorbills (Alca torda) group of three on cliff face, Farne Islands, Northumberland, England, UK, June
Razorbill (Alca torda), pair on cliff, Farne Islands, Northumberland, England, UK, June
Belchers / Peruvian band-tailed gull (Larus belcheri) calling, Paracas National Reserve, Peru
Razorbill (Alca torda) Great Saltee Island, County Wexford, Ireland, June
Northern fulmar (Fulmarus glacialis) nesting pair, Saltee Islands, County Wexford, Republic of Ireland, June