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Mauve stinger (Pelagia noctiluca) at the entrance of a cave, Sula Sgeir, Outer Hebrides / Western Isles, Scotland, July
Atlantic puffin (Fratercula arctica) diving, North Rona, Outer Hebrides / Western Isles, Scotland, UK, July
Mountain hare (Lepus timidus) in winter coat running across snow-covered upland, Scotland, UK. January
Great Skua (Stercorarius skua) silhouetted at sunrise, Fair Isle, Shetland, Scotland, UK, June
Mountain hare (Lepus timidus) in white winter coat, Cairngorms National Park, Scotland, UK, January
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
View over Uath Lochans before sunrise following heavy snowfall, Cairngorms National Park, Scotland, UK, November 2015
European river otter (Lutra lutra) swimming over kelp, Shetland, Scotland, UK, April
Hen harrier (Circus cyaneus) recently fledged chick in flight, Scotland, UK. July
Rock formations on Cul Mor coated in ice. Assynt, Highlands of Scotland, UK, January 2016
Alt Ruadh river flowing through woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, October 2013
Aerial view over Insh Marshes National Nature Reserve, Cairngorms National Park, Scotland, UK, May 2016
Waves crashing on Loch Bad a Ghaill, Assynt, Highlands of Scotland, UK, January 2016
Sparce birch woodland in Glen Etive. Loch Etive, Highlands of Scotland, UK, January 2016
A lone hiker stands against blizzard conditions on An Teallach. Ullapool, Highlands of Scotland, UK, January 2016
First light on Ben Starav ridge in full winter conditions. Glen Etive, Highlands of Scotland, UK, January 2016
Ridge up to Ben Starav in winter. Glen Etive, Highlands of Scotland, UK, January 2016
Sandstone pinnacles and view of lochs below, Stac Pollaidh Inverpolly, Highlands of Scotland, UK, February
Fionn Loch at sunset. Great Wilderness, Highlands of Scotland, UK, January 2016
Devils Point and River Dee at sunrise, in snow, Cairngorms National Park, Highlands of Scotland, UK, January 2016
Grey seal (Halichoerus grypus) pup hauled out on rocky beach, west coast of Scotland, September
Whooper Swan (Cygnus cygnus) with snowy mountains behind, Loch Etive, Agyll and Bute, Scotland, UK, March. Small repro only
Red squirrel (Sciurus vulgaris) on branch in oak woodland with Loch Awe in background. Argyll and Bute, Scotland, UK, June
Red squirrel (Sciurus vulgaris) leaping with nut in mouth. Scotland, UK. Small repro only
Scots Pine (Pinus sylvestris) silhouetted at dawn, Cairngorms National Park, Scotland, UK, October 2013
Osprey (Pandion haliaetus) male wing detail, Cairngorms National Park, Scotland, UK, August
