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Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Brown bear (Ursus arctos) paw seem from under water, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
Wild female Amur leopard (Panthera pardus orientalis) looking up, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Arctic fox (Vulpes lagopus) in winter fur, licking nose, Wrangel Island, Far Eastern Russia, October
Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, June
Polar bear (Ursus martimus) footprints in snow, Wrangel Island, Far Eastern Russia, March
Polar bear (Ursus maritimus) female with young, age one year and a half, resting on the ice, Wrangel island, Far East Russia
Amur leopard (Panthera pardus orientalis) Land of the Leopard National Park, Primorsky Krai, Far East Russia. March
Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, May
Female Brown bear (Ursus arctos) with two cubs, eating fish in lake Kuril, Kronotsky Nature Reserve, Kamchatka, Russia
Female Brown bear (Ursus arctos) with cub playing in lake Kuril, Kronotsky Nature Reserve, Kamchatka, Russia
Siberian tiger (Panthera tigris altaica) at night, taken with remote camera in Land of the Leopard National Park, Far East Russia
Arctic fox (Vulpes lagopus) in summer coat calling, Wrangel Island, Far Eastern Russia, August
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, September
Snowy owl (Bubo scandiacus) in flight, Wrangel Island, Far Eastern Russia, October
Arctic fox (Alopex lagopus) juveniles play fighting, Wrangel Island, Far East Russia, August
Arctic fox (Alopex lagopus) running in snowy landscape with mountains behind, Wrangel Island, Far East Russia, May
Snowy owl (Bubo scandiacus) fledgling flapping wings, Wrangel Island, Far Eastern Russia, August
Polar bear (Ursus maritimus) walking along beach, Wrangel Island, Far Eastern Russia, September
Musk ox (Ovibos moschatus) herd, Wrangel Island, Far Eastern Russia, March
Polar bear (Ursus maritimus) mother with three very young cubs, Wrangel Island, Far Eastern Russia, March
Snow goose (Chen caerulescens caerulescens) brooding chicks, with rusty orange face from iron rich soil in which it forages. Wrangel Island, Far Eastern Russia, June
Polar bear (Ursus maritimus) female with juvenile, age one year and a half, Wrangel Island, Russia
Underwater view of Brown bear (Ursus arctos) paw fishing for Sockeye salmon (Oncorhynchus nerka) with paw outstretched. Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August
Polar bear (Ursus maritimus) adult female with young age one year and a half. Wrangel island, Far East Russia
Raccoon dog (Nycterentes procyonoides) walking across snow, Vladivostok, Primorsky Krai, Far East Russia. November
Amur leopard cat (Prionailurus bengalensis euptilurus) leaping past reed bed, Vladivostok, Primorsky Krai, Far East Russia. February
Arctic fox (Vulpes lagopus) in summer moult stealing Snow goose (Chen caerulescens caerulescens) egg, with Snow goose flapping wings cand calling, Wrangel Island, Far Eastern Russia, June
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Red Foxes (Vulpes vulpes) young play fighting. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, October
Musk ox (Ovibos moschatus) portrait of two standing closely, Wrangel Island, Far Eastern Russia, September
Red fox (Vulpes vulpes) cub biting tail of adult male, Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East. June
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Sika deer (Cervus nippon) doe standing on rocky outcrop overlooking mountain forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. January
Sika deer (Cervus nippon) stag wading through river in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) herd feeding beside pool in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) doe with fawn standing on rocky outcrop overlooking forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Asian black bear (Ursus thibetanus) scent marking tree by rubbing back on it, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it, Land of the Leopard National Park, Russian Far East. Endangered
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Amur leopard (Panthera pardus orientalis) walking past snow-covered trees on mountain slope, Land of the Leopard National Park, Russian Far East. Critically endangered. Taken with remote camera
Sika deer (Cervus nippon) doe walking across snow covered mountain slope, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Snow goose (Chen caerulescens) mobbing Arctic fox (Alopex lagopus) Wrangel Island, Far East Russia, June