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African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA
Domestic dog, Dogo Argentino, three puppies playing with stick, France
Two Red deer (Cervus elaphus) stags, fighting during the rut, Cheshire, UK. July
Greater flamingos (Phoenicopterus ruber roseus) males displaying, lake Nakuru, Kenya
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Dalmatian pelicans (Pelecanus crispus) fighting for fish. Lake Kerkini, Greece. February. Vulnerable species
Two Grey heron (Ardea cinerea) fighting over a fish, Pusztaszer reserve, Hungary. May
Black stork (Ciconia nigra) and Grey heron (Ardea cinerea) Pusztaszer reserve, Hungary
Greenfinch (Carduelis chloris) and Goldfinch (Carduelis carduelis) fighting in snow, Poland. February
African elephant (Loxodonta africa) male in threat display to rival. Floor of the Ngorongoro Crater, Ngorongoro Conservation Area, Tanzania
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
Paradise Riflebird (Ptiloris paradiseus) young males size each other up between practice displays. Australia
Two male Alpine ibex (Capra ibex ibex) fighting in front of the Mer de Glace glacier, with a female watching, Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2
