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RF- Traigh Losgaintir beach and estuary in evening light, North Harris, Western Isles / Outer Hebrides, Scotland, UK. May 2011
Underwater view of tree, with sun rays entering the water, Fibreno Lake nature reserve, Lazio, Italy
Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean
Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Giant Pacific Manta Ray (Manta birostris) portrait with Remora fish (Echeneididae) Roca Partida, Revillagigedos Islands, Mexico. East Pacific Ocean. February
Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Southern stingray (Hypanus americanus) at the surface. Stingray City, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Bait was used for this shot
Round ribbontail ray (Taeniura meyeni) South Ari Atoll, Maldives. Indian Ocean
Ocellated eagle ray (Aetobatus ocellatus), underside, Natal Coast, South Africa
Southern stingray (Hypanus americanus) swimming over ripples on sandbar, Grand Cayman, Cayman Islands
A school of Southern stingrays (Hypanus americanus) on a sand bar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Oceanic manta ray (Mobula birostris) accompanied by three Black jacks (Caranx lugubris) silhouetted, Revillagigedo Islands, Mexico, Pacific Ocean
RF- Shaft of Sunlight in Beech woodland in autumn, Ashridge, Hertfordshire, UK
Wych Elm (Ulmus glabra) dead trees in mist at dawn, Berwickshire, Scotland, July
Group of Spotted eagle rays (Aetobatus narinari) swimming in shallow water, Santa Cruz Island, Galapagos National Park, Pacific Ocean. Small repro only
Reef manta ray (Mobula alfredi) swimming with a Twinspot snapper (Lutjanus bohar), Okinawa, Japan, Pacific Ocean
Reef manta ray (Mobula alfredi) feeding on plankton aggregating in the lights from a boat at night, Ari Atoll, Maldives, Indian Ocean
Group of Clarion angelfish (Holacanthus clarionensis) cleaning an Oceanic manta ray (Mobula birostris) as it swims over the reef, Revillagigedo Islands, Mexico, Pacific Ocean
Oceanic manta ray (Mobula birostris) rolling over on its back as it is cleaned by Clarion angelfish (Holacanthus clarionensis), Revillagigedo Islands, Mexico, Pacific Ocean
Two Oceanic manta rays (Mobula birostris) waiting to be cleaned by Clarion angelfish (Holacanthus clarionensis), Revillagigedo Islands, Mexico, Pacific Ocean
Large school of Pacific cownose rays / Golden cownose rays (Rhinoptera steindachneri), Sea of Cortez, Baja California, Mexico
Manta ray (Mobula alfredi) feeding on plankton aggregated in the lights from a boat at night, North Ari Atoll, Maldives, Indian Ocean
Ocellated eagle ray (Aetobatus ocellatus) searching for food in the sand over the seafloor, Mu Koh Similan National Park, Andaman Sea, Phang-nga, Thailand
RF - Reef manta (Mobula alfredi) swimming in open water with a Remora (Remora remora) swimming below, North Ari Atoll, Maldives, Indian Ocean
Manta ray (Mobula alfredi) with mouth open, feeding on plankton near the surface, Hanifaru Lagoon, Baa Atoll, Maldives, Indian Ocean
Large school of Munks devil rays (Mobula munkiana) aggregating. West Coast of Baja California Peninsula, Mexico. Pacific Ocean
A huge school of Pacific cownose / Golden cownose rays (Rhinoptera steindachneri), Baja California, Sea of Cortez, Mexico
Bluespotted ribbontail ray (Taeniura lymma) searching for food over the coral reef at night, Triton Bay, West Papua, Indonesia, Pacific Ocean
Reef manta ray (Mobula alfredi) feeding, with gills and cephalic lobes visible, Dampier strait, Raja Ampat, West Papua Indonesia, Pacific Ocean
Pink whipray (Pateobatis fai) in shallow water, Moorea, French Polynesia, Pacific Ocean
School of Munks devil ray, pygmy devil ray, (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico
Red stingray (Hemitrygon akajei) with Banded houndshark (Triakis scyllium) group and smaller fish in background. Chiba Prefecture, Honshu, Japan. September
Red stingray (Hemitrygon akajei), four in a row, viewed from below. Chiba Prefecture, Honshu, Japan. October
Longtail stingray (Hypanus longus) on reef, Whitetip reef shark (Triaenodon obesus) group in background. Socorro Island, Mexico
Circular stingaree (Urolophus circularis) above sea floor. Albany, Western Australia. October
Atlantic mobula ray (Mobula hypostoma) group swimming in Caribbean Sea. Isla Mujeres, Quintana Roo, Mexico
Reef manta (Mobula alfredi) visiting a cleaning station on a coral reef with a school of Onespot snappers (Lutjanus monostigma)
Spotted eagle rays (Aetobatus narinari), in mangrove inlet, Elizabeth Bay, Isabela Island, Galapagos
Marbled Stingray (Taeniurops meyeni), Galapagos
A group of large stingrays (Dasyatis americana) swim over sand in shallow water, in a split level photo taken at dawn. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Horned ray (Mobula tarapacana) El Hierro, Canary Islands
Golden Rays (Rhinoptera steindachneri) swimming with Green turtle (Chenolia mydas). Isabela Island. Galapagos
Yellow stingray (Urobatis jamaicensis) on sand seabed by coral reef, The Bahamas
Split level image of a Southern stingray (Dasyatis americana) swimming over a sand bar. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Spotted eagle rays (Aetobatus narinari) through a large window on the wreck of the former mine sweeper C-56 Juan Escutia. Cancun, Mexico
A split level photo of a female southern stingray (Dasyatis americana) swimming over seabed. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Reef manta (Mobula alfredi) female swimming close to a coral reef, while cleaner wrasse (Labroides dimidiatus), tiny by comparison, pick parasites from her lips