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A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Green turtle (Chelonia mydas) from underneath swimming through the surface, Mayotte Island, Comores, Indian Ocean
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
Tiger (Panthera tigris tigris), male portrait, Bandhavgarh, India, February 2013
Common dolphin (Delphinus delphis) surfacing, Atlantic ocean, Portugal, September
RF - Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
A shoal of Golden grey mullet (Liza aurata) swim in the surf breaking on the shore of El Hierro Island. Canary Islands, Spain. East Atlantic Ocean
RF - Maldive anemonefish (Amphiprion nigripes) swimming above their host Magnificent sea anemone (Heteractis magnifica) on coral reef
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
RF- Arabian Oryx (Oryx leucoryx) Dubai Desert Conservation Reserve, Dubai, UAE
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
RF- Great grey owl (Strix nebulosa) in flight. Kuhmo, Finland
Polar bear (Ursus maritimus) walking along coast of Wrangel Island, Far Eastern Russia, October
RF - Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean
Demoiselle crane (Grus virgo or Anthropoides virgo), large flock at their wintering site, Rajasthan, India
Great egret (Ardea alba) in flight, Florida, USA, March
RF- White stork (Ciconia ciconia) in flight, La Serena, Extremadura, Spain. March
RF- White-tailed Sea Eagle (Haliaeetus albicilla) turning in flight, Flatanger, Norway, August
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Portree, Skye, Inner Hebrides, Scotland, UK. June
Pine marten (Martes martes) leaping from branch, orange glow in sky behind from the lights of Inverness. Black Isle, Scotland, UK, November. Photographed by camera trap
Male black rhinoceros (Diceros bicornis) silhouetted against clouds, Masai-Mara Game Reserve, Kenya. March
Ptarmigan (Lagopus mutus) in winter plumage in flight, Utsjoki, Finland, April
Red-crowned Crane (Grus japonensis) in flight, Hokkaido, Japan, February
Greater flamingos (Phoenicopterus ruber) in flight over sand dune, Donana National Park, Andalusia, Spain, March
Honey bee (Apis mellifera) in flight, Surrey, England, July
Silvertip sharks (Carcharhinus albimarginatus) in open water, Revillagigedo islands, Mexico. Pacific Ocean
Three Short-finned pilot whales (Globicephala macrorhynchus) close to the surface against the light, Costa Rica. Pacific ocean
European Herring Gulls (Larus argentatus) in flight at sunset Norway, May
European Herring Gull (Larus argentatus) flock in flight at sunset Norway, May
Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK, January
Curimbata (Prochilodus lineatus) in a clear stream of the Pantanal, with Water hyacinth (Eichhornia sp.), Mato Grosso do Sul, Brazil. Taken for the Freshwater Project
Bottlenose Dolphin (Tursiops truncatus) porpoising, Sado Estuary, Portugal
White naped cranes (Grus vipio) in flight, Kyushu, Japan
Phayers Leaf monkey (Trachypithecus phayrei) Gaoligongshan NP, Yunnan province, China
Cheetah (Acinonyx jubatus) running, Kalahari Desert, Botswana
Swallowtail (Papilio machaon) in flight, Finland, Europe
Snowy owl (Bubo scandiaca) taking off over snow, Canada
Sparrowhawk (Accipiter nisus) flying low over vegetation, Falsterbo Sweden August
Steppe Eagle (Aquila nipalensis) in flight at dawn over misty forest, Czech Republic, November
Common darter dragonflies (Sympetrum striolatum) two females resting on reed, Dunsdon Nature Reserve, Devon, UK. August
Giant Anteater (Myrmecophaga tridactyla) approaching livestock fence, Pantanal. Brazil
Whiskered tern (Chlidonias hybridus), Hungary, June
