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Taiwan / Formosan yellow-throated marten (Martes flavigula chrysospila) Alishan National Recreational Forest, Taiwan
Close up at the dental plates of Three color parrotfish (Scarus tricolor) as it comes in for a bite of hard coral, Fiji
Taiwan yellow tit ( Machlolophus holsti ) Lalashan National Forest Reserve, Taiwan. Endemic
Guanaco (Lama guanicoe) in grassland with Torres del Paine rock towers in background. Torres del Paine National Park, Patagonia, Chile. January 2020
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Coarse chameleon (Trioceros rudis) profile, Volcanoes National Park, Rwanda
A portrait of a clown triggerfish (Balistoides conspicillum) on a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Misool Marine Protected Area. Ceram Sea. Tropical West Pacific Ocean
A male pajama cardinalfish (Sphaeramia nematoptera) incubating eggs in his mouth (not clearly visible). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Apple hawk-moth (Langia zenzeroides formosana) Nzaus Mamei Jianshih Village, Hsssinchu County, Taiwan, Controlled conditions
Stellers sea eagle, (Haliaeetus pelagicus), on sea pack ice, Okhotsk Sea, Rausu, Hokkaido, Japan
Palm Cockatoo (Probosciger aterrimus) feeding on Beach Almond (Terminalia catappa) Lockhart River Foreshore, Lockhart River, Cape York Peninsula, Australia
TemminckAśs tragopan (Tragopan temminckii) male during courtship displayTemminck's tragopan (Tragopan temminckii) male during courtship display, showing red-blue lappets. Tibet, China
Arabian gazelle (Gazella gazella), Jaluni, Oman, May
Whooper swan (Cygnus cygnus) in snow. Jyvaskyla, Central Finland. March
Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March
Siberian tiger (Panthera tigris altaica) at night, taken with remote camera in Land of the Leopard National Park, Far East Russia
Brown skua (Stercorarius antarcticus), Ronge Island, Antarctica
Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
Fire-tailed sunbird (Aethopyga ignicauda) perched amongst blossom. North Sikkim, India. April
RF - Giant anteater (Myrmecophaga tridactyla) walking on ranch, with cattle in the background. Los Llanos, Unarmas Reserve
Black rhino (Diceros bicornis) with very long horn, Lewa Wildlife Conservancy, Laikipia, Kenya. October
Brimstone butterfly (Gonepteryx rhamni) resting among foliage, Meeth Quarry, Devon, UK. June
Shoebill stork (Balaeniceps rex) eating a fish in the swamps of Mabamba, Lake Victoria, Uganda
Bactrian camel (Camelus bactrianus) male silhouetted at sunset, living in the wild but owned by a camel herdsman, Turpan Basin, Gobi Desert, Xinjiang, China
Walia ibex (Capra walie) male with Giant lobelia (Lobelia rhynchopetalum) at around 4, 400 meters near Bwahit Pass, Simien Mountains National Park, Amhara, Ethiopia, September
Red fox (Vulpes vulpes) fox waking up from sleep in den in deep snow, stretching and yawning, Kamchatka, Far east Russia, November
Heath fritillary butterfly (Melitaea athalia) on daisy flower, Pyrenees National Park, France, June
Nazca booby (Sula granti) on the coast of Wolf Island, Galapagos
Galapagos dove (Zenadia galapagensis) on head of Galapagos giant tortoise, (Chelonoidis darwini) Santiago Highlands, Galapagos
Feral domestic cattle, Puerto Fragata, Isabela Island, Galapagos
Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire
Oriental Magpie Robin (Copsychus saularis) female, Whitefield, Bangalore, India, March
American crocodile (Crocodylus acutus), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
Great anglehead lizard (Gonocephalus grandis) male, Tioman Island, Malaysia
Hippopotamus (Hippopotamus Amphibius) with back covered in water hyacinth on back after leaving water. Mana Pools National Park Zimbabwe
Tibetan gazelle or Goa, (Procapra picticaudata) leaping, in mid air. Dulan Nature Reserve, Tibetan Plateau, Qinghai, China. October
Chital (Axis axis) female leaping. Tadoba Andhari Tiger Reserve / Tadoba National Park, Maharashtra, India
Pine grosbeak (Pinicola enucleator), perched by berries in Rowan tree, Finland, January
Wood warbler (Phylloscopus sibilatrix) amongst willow catkins, Finland, May
Common loon (Gavia immer), on a lake, Michigan, USA. June
Garibaldi (Hypsypops rubicundus) in forest of giant kelp, (Macrocystis pyrifera) off Santa Barbara Island, California, USA
Moose (Alces alces) bull, portrait. Grand Teton National Park, Wyoming, USA. September
Carters semaphore gecko (Pristurus carteri) standing on rocky ground. Oman, June
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
African penguin (Spheniscus demersus) standing on rock at dusk, with clustered penguins in background. South Africa. November
Greater flamingo (Phoenicopterus roseus) pair at rest in water, Cape Town, South Africa, July
Atlantic puffin (Fratercula arctica) swimming on the ocean surface, Svalbard, Norway, June