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Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, September
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, July
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Gannet (Morus bassanus) fluffy chick with its parent. Shetland Islands, Scotland, UK, August
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Male Blue-footed booby (Sula nebouxii) walking on sand, San Cristobal Island, Galapagos, South America
Great frigatebird (Fregata minor) pair, male on left. Galapagos, Ecuador
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Grey heron (Ardea cinerea) squabbling with Little cormorant (Microcarbo / Phalacrocorax niger) Keoladeo national park, Bharatpur, Rajasthan, India
Northern gannet (Morus bassanus) in flight over colony, Quebec, Canada, August
Gannet (Sula bassana) diving, July UK Shetland
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Little pied cormorant (Microcarbo melanoleucos) perched on branch protruding out of water. Eden NSW, Australia
Red-footed booby (Sula sula), brown morph, Genovesa Island, Galapagos Islands
Nazca booby (Sula granti) on the coast of Wolf Island, Galapagos
Magnificent frigatebirds (Fregata magnificens) diving for fish prey, Borrero Bay, Santa Cruz Island, Galapagos
Magnificent frigatebird (Fregata magnificens) attacking Blue footed booby (Sula nebouxii) feeding chick. This bird is trying to steal the food from the parent. Santa Cruz Island, Galapagos
Blue-footed booby (Sula nebouxii) with chicks begging for food, Santa Cruz Island, Galapagos, Ecuador
Gannet (Morus bassanus) Shetland, England, UK. July
Magnificent frigatebird (Fregata magnificens) in flight over sea between Isabela Island and Fernandina Island, Galapagos
Red-footed booby (Sula sula) perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair on rocky shore at dawn. Punta Suarez, Espanola Island, Galapagos. May 2015
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
Nazca booby (Sula granti) portrait, Galapagos, Ecuador
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Aerial view of nesting Gannets (Morus bassanus) and swirling seas, Hermaness, Shetland, Scotland, UK, October
