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Tompot blenny (Parablennius gattorugine) in bright summer mating colours, peering out from his home in a discarded tube, where he was guarding his eggs (not visible in this photo), Swanage Pier
Splendid leaf frog (Cruziohyla calcarifer) La Selva Field Station, Costa Rica
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Flock of ducks silhouetted against sunset flying over field in winter in the Uitkerkse Polder nature reserve near Blankenberge, West Flanders, Belgium. February 2020
Robin (Erithacus rubecula) at RSPB Leighton Moss Bird Reserve, Silverdale, Lancashire, England, UK. February 2016
Glanville fritillary butterfly (Melitaea cinxia) on blade of grass, Pollino National Park, Basilicata, Italy, May 2009
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
The River Thames and Houses of Parliament at Westminster at sunset with gull, UK, November 2005
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
Deepsea cirrate octopod {Stauroteuthis syrtensis} 830m, Gulf of Maine, Atlantic
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Great Staple Tor in evening light, Dartmoor National Park, Devon, UK
Common toadflax (Linaria vulgaris), orange nectar guides on lower lip and long spur containing nectar. Surrey, England, UK. Focus stacked
Nudibranch (Flabellina nobilis) photographed in the field aquarium in Gulen, Norway. North East Atlantic Ocean
Close up of Garibaldi fish face {Hypsypops rubicunda}, Channel Islands, California, USA
River Brue at dawn near Glastonbury on the Somerset Levels, Somerset, UK, November 2010
Ultramel Okeetee corn snake, with recently laid eggs, an interspecies hybrid between a Corn snake, (Pantherophis guttatus), and a Grey rat snake (Pantherophis spiloides)
Golden mantella frog (Mantella aurantiaca) with freshly laid eggs on wet mossy ground
Corn snake (Pantherophis guttatus), female with recently laid eggs, captive, native to Eastern United States
Autumn quaking aspen grove (Populus tremuloides) with trunks scarred by browsing elk (Cervus canadensis). Below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
Autumn quaking aspen trees (Populus tremuloides) below Turret Ridge, Uncompahgre National Forest, Colorado, USA. October
Crown imperial lily (Fritillaria imperialis), six nectaries at base of stamens and stigma. In cultivation, Native to southeast Turkey and western Himalayas
Sun rising through mist behind Salisbury Cathedral viewed across Harnham Water Meadows. Salisbury Wiltshire, England. March 2009
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Marsh fritillary (Euphydryas aurinia) on fern. South of Casteil, Pyrenees Orientales, south west France. May
Saturniid moth (Eacles ormondei), female camouflaged in leaf litter. Izabal, Guatemala
Three spot damselfish (Dascyllus trimaculatus) with Sea anemone home, Yap, Micronesia
Monarch butterfly (Danaus plexippus), in wintering from November to March in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve, Mexico
Polar bear (Ursus maritimus) silhouetted against setting sun, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Orange tree (Citrus sinensis) flowers and fruit, Crete, Greece, April 2009
Common beech (Fagus sylvatica) woodland in autumn, Cairngorms National Park, Scotland, UK, October
Mixed woodland of ash alder, oak beech in autumn colours. Kinnoull Hill Woodland Park, Perthshire, Scotland, Novermber 2011
Flock of White ibis (Eudocimus albus) in breeding plumage, at rookery on water sFlock of White ibis (Eudocimus albus) in breeding plumage, at rookery on waters edge of the mangrove-covered island, Tampa Bay, Florida, USA, March
A misty morning over a mixed woodland in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011. Highly commended in the Wild Woods Category of the British Wildlife Photography
Grey heron (Ardea cinerea) adult in breeding plumage, close-up of head and colourful orange beak, backlit, Berlin Zoological Garden, Berlin Zoo, Germany. February
Fringed leaf frog (Cruziohyla craspedopus) on branch, Yasuni National Park, Orellana, Ecuador
Laguna Miluni, contaminated with mine effluent and with a low level due to drought. This reservoir is fed by glacial meltwater from the Andean peak, Huayna Potosi, Bolivian Andes. October 2015
Red deer (Cervus elaphus) stag bellowing in mist at sunrise, rutting season, Bushy Park, London, UK, October
Kingfisher (Alcedo atthis) male passing fish to female spring courtship behaviour, Hertfordshire, England, UK, March. Sequence 4 of 6
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
Detail of Plumose anemone (Metridium senile) in its retracted state at slack water when it is not feeding. Plymouth Sound, Devon, England, UK, English Channel, July
Arabian Oryx (Oryx leucoryx) at sunset. Previously extinct in the wild, their conservation status is now stable thanks to conservation efforts.This male is part of a reintroduction program in Dubai
Guianan Cock-of-the-rock (Rupicola rupicola), during courtship display, Guianan Shield, Suriname
Emerald tree Boa (Corallus batesii) close up portrait of juvenile whilst hanging on a branch. Tambopata, Madre de Dios, Peru
Comma butterfly (Polygonia C-album) feeding on ripe blackberries, Dorset, UK, August
