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American crocodile (Crocodylus acutus) with open mouth amongst Seagrass (Alismatales). Gardens of the Queen National Park, Cuba
Giant panda (Ailuropoda melanoleuca) dusted with snow eating Bamboo. In natural enclosure, Panda Breeding Centre, Wolong, Sichuan Province, China. February 2008
Flightless cormorant (Phalacrocorax harrisi) underwater, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador, April
Galapagos land iguana (Conolophus subcristatus) portrait with mouth open, Galapagos
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Black skimmer (Rynchops niger) foraging along surf line. Tierra Verde, Florida, USA
Hippopotamus (Hippopotamus amphibius) in pool with mouth open. Mana Pools National Park, Zimbabwe
Antarctic fur seal (Arctocephalus gazella), mother warning off curious juvenile. Southern elephant seal (Mirounga leonina) in background. Gold Harbour, South Georgia. October
Tokay gecko (Gekko gecko) Pat Sin Leng Country Park, New Territory, Hong Kong, China
Mandrill (Mandrillus sphinx) male showing aggression, captive
Greylag goose (Anser anser) taking flight from lake. Southern Norway. March
Grasshopper Warbler (Locustella naevia) singing, Joensuu, Finland, May
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Male Yellow boxfish (Ostracion cubicus) displaying, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Female Jaguar (Panthera onca) playing with her cub, captive, occurs in Southern and Central America
Lion (Panthera leo) male smelling, flehmen response, Masai-Mara Game Reserve, Kenya. Vulnerable species
Red Foxes (Vulpes vulpes) young play fighting. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Wilsons Storm Petrel (Oceanites oceanicus) feeding whilst walking on water Atlantic ocean, Madeira Portugal, August
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Yacare Caiman (Caiman yacare) with mouth open to keep cool, Pantanal, Brazil
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Robin (Erithacus rubecula) singing, Norfolk, England, UK, February
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Konik Wild Horse (Equus ferus caballus) yawning. The Netherlands, November
A Malinois / Belgian Shepherd police dog Mia owned by German police officer and dog handler, playing in the snow. Germany
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Black blotched pufferfish (Diodon liturosus) inflated with water at night as defensive behaviour, Sipadan Island, Borneo, Malaysia
Red fox (Vulpes vulpes) vixen yawning in the morning light (captive). Lifton, Devon, UK, May
Mixed Breed dog running on grass, towards camera
Head portrait of Sumatran tiger (Panthera tigris sumatrae) with face half cast in shadow, with aggressive expression, captive
Red howler monkey (Alouatta seniculus) howling, captive
Angler fish (Melanocetus murrayi) Mid-Atlantic Ridge, North Atlantic Ocean
Maine Coon female cat, Serafin, in fierce defensive posture
Spectacled caiman (Caiman crocodilus) Rio BaiAia Bonita, Bonito, Mato Grosso do SulSpectacled caiman (Caiman crocodilus) Rio Baiia Bonita, Bonito, Mato Grosso do Sul, Brazil
Asiatic lion cub (Panthera leo persica) looking up into tree, possibly at a bird, Gir Forest NP, Gujarat, India
Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November
RF- Male Bengal Tiger (Panthera tigris tigris) face portrait in undergrowth, Kanha National Park, Madhya Pradesh, India. Endangered species
Eastern Hermann's tortoise (Testudo hermanni boettgeri) hatching from egg, mouth open. Captive, occurs in South East Europe
Robin (Erithacus rubecula) male singing, perched on branch. Joutsa, Finland. April
Cunner (Tautogolabrus adspersus) pair in Eelgrass (Zostera marina) bed. Newfoundland, Canada. September
Ring-tailed lemur (Lemur catta) suckling baby. Berenty Reserve, Madagascar
