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Brown Bear (Ursus arctos) portrait in water. Finland, Europe, June
Bison (Bison bison) in snowstorm. Yellowstone National Park, USA, February
Arctic Fox (Vulpes lagopus) portrait in winter coat. Norway, Captive, March
African Lion (Panthera leo) young male at sunrise, Etosha National Park, Namibia
Common zebra (Equus quagga) foal in dusty sunset, Etosha National Park, Namibia Photo by Sharon Heald
Red fox (Vulpes vulpes) sniffing beech tree trunk, Black Forest, Germany, Winner of Fritz Polking portfolio prize, GDT 2011 competition
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Quinling Golden snub nosed Monkey (Rhinopitecus roxellana qinligensis), infant sitting in a tree. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
A rare East Bulgarian stallion cantering at the Kabiuk National Stud, Shumen, Bulgaria, February 2012
Fallow deer (Dama dama) in woodland clearing at dawn, Cheshire, UK November
DELETE DUPLICATE - Fallow deer (Dama dama) in woodland clearing, Cheshire, UK August
RF- Secretary blenny (Acanthemblemaria maria) peering from hole in massive Starlet coral (Siderastrea siderea). East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Northern raccoon (Procyon lotor) young at night climbing live Oak tree (Quercus virginiana) Dinero, Lake Corpus Christi, South Texas, USA
Long-jawed longhorn beetle (Trachyderes mandibularis) close up head portrait, Dinero, Lake Corpus Christi, South Texas, USA
Domestic dog, Saluki / Arabian Hound / Gazelle Hound / Persian Greyhound, studio portrait, France
Domestic rabbit, French Lop, Blue, young male, 6 months, studio portrait
Brinicles (brine icicles or ice stalactites) forming under the ice, growing towards the sea floor, Ross Sea, McMurdo Sound, Antarctica
Tibetan terrier, sitting, portrait
Looking down at Chubb (Leuciscus cephalus) at surface with reflection of bridge, Rhine River, Switzerland, 1993. No release available
Grey angelfish (Pomacanthus arcuatus) portrait, Georgetown, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Schoolmaster snapper (Lutjanus apodus) above a coral reef, Georgetown, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Briard / Berger de Brie, head portrait with plaited hair between eyes
Rothschilds giraffe (Giraffa camelopardalis rothschildi) close up of young calf skin pattern, captive
Grevy's zebra (Equus grevyi) close up, captive
Red fox (Vulpes vulpes) close-up of half of face, captive
Black bear (ursus americanus) cub standing on hind legs, Yellowstone National Park, USA, June
Grizzly Bear (Ursus arctos horribilis) male in water. Coastal Katmai National Park, southwest Alaska, USA, August
Blue Shark (Prionace glauca) near sea surface and divers in background. Santa Maria, Azores, September
Demoiselle crane (Anthropoides virgo) portrait, captive
Common buckeye butterfly (Junonia coenia) feeding on Ironweed (Vernonia altissima) flower, North Guilford, Connecticut, USA, August
Leatherback Turtle (Dermochelys coriacea) returning to the sea after egg laying. Cayenne, French Guiana, June
Ocellated eagle ray (Aetobatus ocellatus), underside, Natal Coast, South Africa
Black blotched pufferfish (Diodon liturosus) inflated with water at night as defensive behaviour, Sipadan Island, Borneo, Malaysia
Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Red Fox (Vulpes vulpes) vixen on a misty day in woodland, Black Forest, Germany, July. Runner-up in Animal Portraits category, Wildlife Photographer of the Year competition 2011
American Cocker spaniel, USA
Tabby kitten, 5 months, standing up with raised paws
Yorkshire Terrier, lying with chin on the floor
Portrait of a Birman-cross cat
Portrait of a Beagle puppy
Colourpoint kitten stretching
Chocolate Labrador puppy, 3 months, lying
Spotted Hyaena (Crocuta crocuta) rolling in Devils Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January
Portrait of Waglers / Temple Pitviper (Tropidolaemus wagleri) showing the thermo-receptive pit organ between the eye and mouth. Controlled conditions. Mindanao, Philippines, February
A male Spiny Seahorse (Hippocampus guttulatus) hides in seagrass. Etang De Thau (Thau Lagoon), Montpellier, France, Europe, September
West Highland White Terrier leaping