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Chimpanzee female (Pan troglodytes schweinfurthii) sleeping in a nest built in a tree, Kibale National Park, Uganda, January
Female Uganda kob (Kobus kob thomasi) suckling calf. Murchisson Falls National Park, Uganda. February
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Coalition of two African male lions (Panthera leo) resting on the banks of the Luangwa river, dry season, South Luangwa National Park, Zambia
Hawthorn tree and granite outcrop at Holwell tor, sunrise and mist, Dartmoor National Park, Devon, UK. October 2018
Great Mis Tor covered in snow, Dartmoor National Park, Devon, England, UK, January 2015
Great Staple Tor at dusk, Dartmoor National Park, Devon, England, UK, January
Landscape view of Great Staple Tor, Dartmoor National Park, Devon, England, UK, January 2012
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
European otter (Lutra lutra) shaking off water, Kiskunsagi National Park, Hungary, January
Forested island on Loch Lomond, Loch Lomond and the Trossachs National Park, Scotland, UK
Mountain Gorilla (Goriila beringei) resting, Volcanoes National Park, Virunga Mountains, Rwanda
Cuban emerald hummingbird (Chlorostilbon ricordii) Guanahacabibes Peninsula National Park, Cuba
Bee hummingbird (Mellisuga helenae) feeding from flower, Guanahacabibes Peninsula National Park, Cuba
Mountain gorilla (Gorilla beringei beringei), dominant silverback in forest clearing. Volcanoes National Park, Rwanda
Muskox (Ovibos moschatus) in tundra with mountains in background, Dovrefjell National Park, Norway. September 2018
RF - American Bison (Bison bison) male in snow covered in frost, standing by dead trees. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January
Cuban crocodiles (Crocodylus rhombifer) photographed on a crocodile farm started by Fidel Castro in the 1960s in Cienaga de Zapata National Park to help save the species. Cuba
Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January
Freshwater pools on Waterfall Creek, at the top of Gunlom Falls. Mary River, Kakadu National Park, Northern Territory, Australia. March 2014
Bamboo lemur (Hapalmur griseus) sitting on tree stump. Andasibe-Mantadia National Park, Madagascar
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Beech woodland near Blackwater Brook. New Forest National Park. November 2014
Corncrake (Crex Crex) calling, with breath vapor in early morning, Kiskunsagi National Park, Hungary
Black-backed jackals (Canis mesomelas) two fighting with Plains / Burchell zebra (Equus burchelli) in foreground, Kalahari Desert, Nxai Pans National Park. Botswana
Spider webs covered in dew, on a flowering Gorse bush, Peak District National Park, UK October
Giant honey bee nest (Apis dorsata) up in a giant Mengaris tree (Koompassia excelsa) dipterocarp forest in Danum Valley, Sabah, Borneo, Malaysia
Kaieteur Falls is the worlds widest single drop waterfall, located on the Potaro river in the Kaieteur National Park, in Essequibo, Guyana, South America
Mud-covered face of bull African buffalo (Syncerus caffer) with a cateract in one eye, on the banks of the Chobe River, Chobe National Park, Botswana
Great egret (Ardea alba) and Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Mountain landscape reflected in lake, Basongcuo National Park, Qinghai-Tibet Plateau, Tibet, November 2011
Group of Great egrets (Ardea alba), Grey heron (Ardea cinerea) and Eurasian spoonbill (Platalea leucorodia) at night with stars and moon. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May
Suspension bridge across the river, entrance to Gunung Mulu National Park UNESCO Natural World Heritage Site, Malaysian Borneo
Landscape modified by the erosion process. Goreme National Park and the Rock Sites of Cappadocia UNESCO World Heritage Site. Turkey. December 2006
Ordesa y Monte Perdido National Park, Huesca, Aragon, Spain, July 2016
Bornean orangutan (Pongo pygmaeus) male, Gunung Palung National Park, Borneo
Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Young ground squirrels (Xerus inauris) fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Misty morning view of Exmoor countryside from Winsford Hill, Exmoor National Park, Somerset, UK. October 2016
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Pristine tropical rainforest and granite cliffs in mist, Morne Seychelles National Park, Mahe Island, Republic of Seychelles
Egyptian goose (Alopochen aegyptiaca) pair amongst Devil s-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia, March
Scimitar-horned oryx (Oryx dammah) captive in enclosure of Souss Massa National Park, Morocco. Extinct in the Wild
Hippopotamus (Hippopotamus Amphibius) territorial aggressive behavior, Chobe National Park, Botswana
Sea apple (Pseudocolochirus violaceus) Komodo National Park, Indonesia
