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African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012
Red kite (Milvus milvus) in flight, Gigrin Farm, Mid Wales, UK, March. Non-ex
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Weasel (Mustela erminea) in winter coat, running through deep snow, Upper Bavaria, Germany, Europe. February
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
Cairns birdwing butterfly (Ornithoptera priamus) male on fern leaf and Blue Mountain Swallowtail butterfly in flight, Atherton Tablelands, Queensland, Australia, October
Lion (Panthera leo) male shaking his head after rain, Masai-Mara Game Reserve, Kenya
RF -Three colourful Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Red-and-green macaw (Ara chloropterus) in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Mountain hare (Lepus timidus) running in half summer coat, Peak District, UK April
Caribbean flamingo (Phoenicopterus ruber) about to take off to roosting site, Ria Celestun Biosphere Reserve, Yucatan Peninsula Mexico, January
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Sequence of a Red-and-green macaw or green-winged macaw (Ara chloropterus) coming into land at its nest hole on a cliff. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Brown bear (Ursus arctos) shaking water from its coat, Carpathian Mountains, Romania. April
Weedy seadragon (Phyllopteryx taeniolatus) swimming over seaweeds. Sydney, New South Wales, Australia. Pacific Ocean
School of powderblue surgeonfish (Acanthurus leucosternon) feeding, Laamu Atoll, Maldives. Indian Ocean
Firefly (Lamprohiza splendidula) light trails in woodland at night, multiple exposure. The Netherlands. June
Munks mobula ray / Devilray (Mobula munkiana) leaping out of the water, Sea of Cortez, Gulf of California, Baja California, Mexico, February
Jaguar (Panthera onca) male and female four month old cubs playing, native to Southern and Central America, captive
Ice in flowing river water. Scotland, UK, February
Fireflies {Lampyridae} in rainforest, Singapore
African elephant charging (Loxodonta africana) female with young calf, Okavango Delta, Botswana
Two Cheetah (Acinonyx jubatus) chasing Springbok (Antidorcas marsupialis) Kgalagadi, South Africa
Bull sharks (Carcharhinus leucas) in Beqa (Benga) Lagoon, a reserve where sharks are fed as part of a commercial diving operation. Viti Levu, Fiji, South Pacific
Spotted reed frog (Hyperolius pucticulatus) tadpoles, captive, endemic to Tanzania
Pair of Blesbok (Damaliscus pygargus) galloping across the open flood plains of the Bushmans River, Kariega Game Reserve, South Africa. Overall winner of the GDT Competition 2014
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
Vehicle tracks in field of ripe Barley, farmland, late evening light, near Putford, Devon, UK. August 2012
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
Puffins (Fratercula arctica) pair take off together to go fishing. Farne Islands, Northumberland, UK, May. Non-ex
African lion (Panthera leo) male shaking water off mane after rain, Masai-Mara Game Reserve, Kenya. September
Landscape view of the Devil's Jaw, Ersfjord, Senja, Norway, February
Southern right whale (Eubalaena Australis) tail fluke breaching the ocean surface against a golden sunset, Peninsula Valdes, Patagonia, Argentina, Atlantic Ocean
Puma (Puma concolor) female, running sequence in deep snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile. Composite of five separate images
White rhinoceros (Ceratotherium simum) charging, Mala Mala Game Reserve, South Africa
Spinner dolphin (Stenella longirostris) juvenile surfacing next to its mother, Sri Lanka
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal
Kerry bog pony, stallion, a rare breed, running, County Kerry, Republic of Ireland. April
Large school of juvenile Fusilier (Caesionidae) fish swimming in water column, Andaman Sea, Mu Koh Similan National Park, Phang-nga, Thailand