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Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef. Whilst a Predatory bandcheek wrasse (Oxycheilinus digrammus)
Poppies (Papaver rhoeas) and cornflowers (Centaurea cyanus) near Castellucio di Norcia, Umbria, Italy, July
Black and white Border collie bitch with black and white tuxedo kitten age 10 weeks
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
Tabby kitten, Picasso, 9 weeks, head to head with Yellow labrador puppy, 8 weeks
Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Bream (Abramis brama) and European chub (Leuciscus / Squalius cephalus) shoal in a calm part of the Rhine, Schaffhausen, Switzerland, February 2009 WWE OUTDOOR EXHIBITION
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern
Baby rabbit with a Roborovski Hamster (Phodopus roborovskii) sitting on its head
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Cape buffalo (Syncerus caffer caffer) portrait, Red-billed oxpeckers (Buphagus erythrorhynchus) removing parasites. Nakuru, Kenya
Orca (Orcinus orca) with South American sealion (Otaria flavescens) in mouth, beaching itself on shore. Punta Norte, Valdez Peninsula, Argentina. April
Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico
Red-headed woodpeckers (Melanerpes erythrocephalus), pair fighting with European Starling (Sturnus vulgaris) that is trying to take over the woodpeckers nest hole, Montezuma National Wildlife Refuge
Sleepy ginger kitten and lionhead cross rabbit
Peruvian Tarantula (Pamphobeteus sp. ) adult, walking over Humming Frog (ChiasmocleisPeruvian Tarantula (Pamphobeteus sp.) adult, walking over Humming Frog (Chiasmocleis royi) without preying on it. Los Amigos Biological Station, Madre de Dios, Amazonia, Peru
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Nudibranchs (Nembrotha chamberlaini) feeding on Tunicates (sea squirt) on a coral reef. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean
Various species of moth from tropical rainforest, Danum Valley, Sabah, Borneo. Digital composite
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Anemone prawn (Periclimenes sagittifer) on Snakelocks anemone (Anemonia viridis) Jersey, British Channel Islands, June
Gorgonian / Sea whip coral (Ellisella ceratophyta) with Ring tailed cardinalfish (Apogon aureus) swimming in front, West Papua, Indonesia
Coral reef with soft corals (Dendronephthya sp. ), encrusting sponges and Pygmy sweepersCoral reef with soft corals (Dendronephthya sp.), encrusting sponges and Pygmy sweepers (Parapriacanthus ransonetti) Similan Islands, Andaman Sea, Thailand
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha ab Mahmud, Sinai, Egypt. Red Sea
Cheetah (Acinonyx jubatus) hunting Springbok (Antidorcas marsupialis) trying to trip up the prey, Kgalagadi, South Africa
Indian gazelle (Chinkara) (Gazella bennetti) with flock of Common pigeons, Lohawat, Rajasthan, India
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Red Howler Monkey (Alouatta seniculus) hanging by tail with non-prehensile tailed Peruvian red uakari monkey (Cacajao calvus ucayalii)
Brush footed butterfly (Lycorea sp. ) caterpillar with parasitic wasp cocoons on silk threadsBrush footed butterfly (Lycorea sp.) caterpillar with parasitic wasp cocoons on silk threads, Yasuni National Park, Orellana, Ecuador
Tropical rainforest invertebrates, Sabah, Borneo. Digital composite
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
Rudbeckia fulgida var. sullivantii Goldsturm in garden with Michaelmas daisies
Polar bear (Ursus maritimus) hunting Seal on sea ice. Svalbard, Norway, July 2018
European mantis (Mantis religiosa) hunting swarming Pale burrower mayfly (Ephoron virgo), mayflies recently hatched. Tudela, La Ribera de Navarra, Navarre, Spain. August
Orca (Orcinus orca) hunting South American sealion (Otaria flavescens) close to shore. Punta Norte, Valdez Peninsula, Argentina. April
Butterflies and day flying moths feed from Loofah flower (Luffa cylindrica) and pick up pollen on their legs, underside of thorax and antennae, South Sichuan Bamboo Sea, China