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Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Oystercatchers (Haematopus ostralegus) arriving to roost at Snettisham on the Wash. Norfolk, January
Pink-footed Geese (Anser brachyrhynchus) in flight at dusk. The Wash. Norfolk, January
Flock of Knot (Calidris canuta) over sea with wind turbine. Liverpool Bay, UK, December
Five images of schooling Raccoon Butterflyfish (Chaetodon lunula) digitally combined for this panorama photograph. Lanai, Hawaii
Flock of Oystercatchers (Haematopus ostralegus) and Knot (Calidris canuta) in flight and on tidal shallows. The Wash Estuary, Norfolk, September
Gannet (Morus bassanus) breeding colony, long exposure showing flight trails. Hermaness National Nature Reserve, Shetland, Scotland, July
Flock of Knot (Calidris canuta) taking flight. Snettisham, UK, January
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Row of Starlings (Sturnus vulgaris) perched on wire. Marais Breton, Vendee, French Atlantic Coast, November