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Whooper swan (Cygnus cygnus) pair on loch in heart shape, Cairngorms National Park, Scotland, UK, November
Five-spot burnet moths {Zygaena trifolii} mating on Pyramidal orchid {Anacamptis pyramidalis} Cornwall, UK
Strawberry poison frog (Oophaga pumilio) pair mating, Sarapiqui, Heredia, Costa Rica
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Four-spot orb weaver spider (Araneus quadratus) male approaching larger female to mate. Peak District National Park, Derbyshire, UK. October
Emerald damselflies (Lestes sponsa) mating, with male above. Peak District National Park, Derbyshire, UK. September
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Two male Common cuttlefish (Sepia officinalis) compete for a female (on right) during courtship, Torbay, Devon, UK, May. 2020VISION Exhibition. 2020VISION Book Plate
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Greater flamingos (Phoenicopterus roseus) mating, in lagoon, Pont Du Gau, Camargue, France, April 2009
Lemur leaf frog (Hylomantis lemur) mating pair, captive, Critically endangered, from Central America. Please credit Michael Kern / Zoo Atlanta
Masai Giraffe (Giraffa camelopardalis tippelskirchi) male and female (back). Masai-Mara Game Reserve, Kenya
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Western grebe (Aechmophorus occidentalis) pair swimming, with reflections, Bolsa Chica Ecological Reserve, California, USA, February
Forester kangaroo (Macropus giganteus tasmaniensis) family group, male, female and large joey, Tasmania, Australia, February. Subspecies of Eastern grey kangaroo
Pipevine Swallowtail Butterfly (Battus philenor) pair mating. Sinton, Corpus Christi, Coastal Bend, Texas, USA, June
A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)
Goldies Bird of Paradise (Paradisaea decora) male displaying to female at lek, Fergusson Island, Milne Bay Province, Papua New Guinea. Near-threatened
Erect-crested penguins (Eudyptes sclateri) pair on coast, Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Mating pair of King Penguins (Aptenodytes patagonicus) on Gold Beach, South Georgia Island, Sub Antarctica
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
Waved albatross (Phoebastria irrorata) courting pair mutually grooming. Punta Cevallos, Espanola (Hood) Island, Galapagos, Ecuador, May
Silky sifaka (Propithecus candidus) pair in tree, Marojejy National Park, Madagascar, Endangered species
Dusky dolphins (Lagenorhynchus obscurus) courting pair leaping from the sea, Kaikoura, Canterbury, New Zealand, January
Humpback whale (Megaptera novaeangliae australis) courtship, with the female above showing her abdomen and the male below. Vava'u, Tonga. Pacific Ocean
Banded demoiselle dragonfly (Calopteryx splendens) male and female silhouetted, resting on reeds. Lower Tamar Lakes, Cornwall, England, UK. June
RF - Puffins (Fratercula arctica) pair, resting on cliffside among flowering Sea thrift (Armeria maritima), Great Saltee Island, County Wexford, Republic of Ireland. May
Pair of Red-eyed damselfly (Erythromma najas) perched on grass blade, at sunset, Poland. June
Banded demoiselle (Calopteryx splendens), male and female silhouetted on reed, Lower Tamar Lakes, Cornwall, UK. June
Ventral view of Swallowtail butterfly (Papilio machaon), male and female, in flight. The Netherlands. July
Whooper swans (Cygnus cygnus) pair engaged in courtship as snow falls, Myvatn lake, Iceland, March
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
Northern gannet (Morus bassanus) pair, courtship, Heligoland, Germany
Western grey kangaroo (Macropus fuliginosus) pair, male standing and female lying down, Kangaroo Island, Australia
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October
Pair of Waved albatrosses (Phoebastria irrorata) billing as greeting. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Pair of Swallow-tailed gulls (Creagrus furcatus) at nest, one incubating. Genovesa Island, Galapagos Islands, Ecuador
Pair of Evening grosbeaks (Coccothraustes vespertinus), in winter plumage, perched on moss-covered log. New York, USA. October. Digitally retouched; distractions removed. Topaz AI Sharpen applied
Humpback whales (Megaptera novaeangliae) pair engaging in courtship behavior, dancing for hours, off Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) male and female engaging in courtship, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) male and female pair courting, Tubuai, French Polynesia, Pacific Ocean
Magnificent frigatebirds (Fregata magnificens) pair in courtship display, male with gular sac inflated, North Seymour Island, Galapagos National Park, Galapagos Islands
Pale-mandibled aracaris (Pteroglossus erythropygius) pair, at entrance to nest hole in tree, Choco rainforest, Los Bancos, Pichincha, Ecuador
Common ostriches (Struthio camelus), male and female, standing at waterhole in front of Grant's zebras (Equus quagga boehmi). Tsavo East National Park, Kenya
Pair of Coleman shrimps (Periclimenes colemani) living in a fire urchin (Asthenosoma varium). The female is the larger shrimp
Pair of Broadclub cuttlefish (Sepia latimanus) mating in a coral reef, Kagoshima Prefecture, Japan. Having succeeded in gaining the female's acceptance
Pair of Siberian flying squirrel (Pteromys volans orii) sharing nest in tree hole in preparation for reproductive season. Hokkaido, Japan. March