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Baltimore oriole (Icterus galbula) female perched on Berlandiers fiddlewood (Citharexylum berlandieri). South Padre Island, Texas, USA. April
RF - Common darter dragonfly (Sympetrum striolatum) resting on Foxglove (Digitalis purpurea) with dew droplets on wings. Cornwall, England, UK. June
Bunch-flowered narcissus (Narcissus tazetta) flower head, growing on limestone, Umbria, Italy. February
Scalesia pedunculata forest, Los Gemelos, Highlands, Santa Cruz Island, Galapagos, South America
Lava cactus (Brachycereus nesioticus) growing on bare lava, Fernandina Island, Galapagos, South America
Snakelocks anemone (Anemonia viridis) growing on a stalk of Eelgrass (Zostera marina) on the seabed, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Large flower of the parasitic plant Rafflesia pricei, growing in rainforest, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
Garden lupin (Lupinus polyphyllus) Terwick Church Fields, Sussex, UK. June
Common kingfisher (Alcedo atthis) perched on a bulrush, Devpm, UK, November
Hercules club / Canary Island Spurge (Euphorbia canariensis) in montane habitat, with flat topped mountain ridge in background, Parque Rural de Teno, Tenerife, Canary Islands
Teide wallflower (Erysimum scoparium) endemic to Tenerife, Teide National Park, Canary Islands; Spain, Europe
Canary samphire (Astydamia latifolia) close up of flowers and leaves, Punta del Teno, Parque Rural de Teno, Tenerife, Canary Islands
Great horned owls (Bubo virginianus), adult and chick, on nest in Saguaro cactus (Carnegiea gigantea), Sonoran desert, Arizona
Tawny owl (Strix aluco) perched in Silver Birch tree (Betula pendula), Dumfries and Galloway, Scotland, UK, May
Kreffts gliders (Petarus notatus) two in Eucalypt (Myrtaceae) tree on branch with blossom at night, Toowoomba, Queensland, Australia
Portuguese oak tree (Quercus faginea) covered in moss, Los Alcornocales Natural Park, Andalusia, southern Spain, November
Ancient hawthorn (Crataegus monogyna) with berries and stripped of leaves, Hergest Ridge looking across to Herrock Hill, Herefordshire / Radnorshire border, UK. October
Variable oysterling (Crepidotus variabilis) on dead Field maple (Acer campestre) twig, Herefordshire, England, UK. November, Focus stacked
Downy birch (Betula pubescens) changing to autumn colours. Caledonian pine forest, Glen Strathfarrar, Scottish Highlands. Scotland. October
Great horned owl (Bubo virginianus) adult and chick in nest in Saguaro cactus (Carnegiea gigantea) Sonoran Desert, Arizona, USA
RF - Split level photo of mangrove scenery, with hard corals (including Goniopora sp. Porites sp.) growing below Red mangrove tree (Rhizophora sp.). Raja Ampat, West Papua, Indonesia
Tench (Tinca tinca) during spawning period amongst European watermilfoil (Myriophyllum spicatum), Sion, Valais canton, southwestern Switzerland
Elephant foot yam (Amorphophallus paeoniifolius) one of largest flowers in the world. Blooming in Papillote Garden, Dominica, West Indies
Termite hill and acacia tree, Shaba National Reserve, North Kenya
Euphorbia tree near Solio escarpment, Laikipia County, Kenya
Caralluma socotrana in blossom, near lake Magadi, Kenya
Palmers globemallow (Sphaeralcea palmeri) and view towards Toro Islet and Guadalupe Island, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, September
Rosemary beetle (Chrysolina americana) feeding on Lavender (Lavandula angustifolia) in garden Cheshire, UK, June
RF - Rogie Falls and autumn woodland with Common oak or Pedunculate oak (Quercus robur). Caledonian forest, Reilig Glen, Scottish Highlands. Scotland. October
Red mangrove (Rhizophora mangle) propagules / plantlets which become fully mature plants before dropping off the parent tree to drift away and establish a new tree. Bahamas
Queen conch (Aliger gigas) in a seagrass (Thalassia testudinum) meadow, Bahamas
Dune pansies / Seaside pansies (Viola tricolor curtisii) flowering on coastal sand dunes, Merthyr Mawr Warren NNR, Glamorgan, Wales, UK, September
Giant Cordon cacti (Echinopsis atacamensis) on Incahuasi Island in Salar de Uyuni salt flats, Bolivia. March
English oak tree (Quercus robur) under stars of the Milk Way, Brecon Beacons National Park, International Dark Sky Preserve, Monmouthshire, Wales, UK
Madonna sun orchid (Thelymitra silena). Tasmania, Australia. October
Gunns tree orchid (Sarcochilus australis). Tasmania, Australia. November
Honeybush (Richea scoparia). Tasmania, Australia. December
Chamomile sunray (Rhodanthe anthemoides). Tasmania, Australia. January
Southern storksbill (Pelargonium australe). Tasmania, Australia. December
Mountain needlebush (Hakea lissosperma). Tasmania, Australia. November
Bordered patch butterfly (Chlosyne lacinia) on Indian blanket (Gaillardia pulchella). Hill Country, Texas, USA
Common spotted orchid and Southern marsh orchid hybrid (Dactyloriza x grandis) in flower. Leicestershire, England, UK. Focus stacked
Common oak (Quercus robur), Sessile oak (Quercus petraea) and hybrid oak in ancient woodland, trees and stones covered in moss. Wistmans Wood, Dartmoor National Park, Devon, England, UK. November 2020
Thrift (Armeria maritima) flowering on cliff top at sunset. Hartland Quay, Devon, England, UK. May 2019
Common spotted orchid (Dactylorhiza fuchsii). Volehouse Moor Devon Wildlife Trust Reserve, England, UK. June
Banyan fig tree (Ficus benjamina), one single individual tree, Banyan garden protected forest, Kenting National Park, Taiwan
Arrowleaf balsamroot (Balsamorhiza sagittata) and lupins (Lupinus sp.), Columbia River Gorge, Oregon, USA. April
Borage (Borago officinalis) watercolour illustration
