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Caribbean reef shark (Carcharhinus perezi) swimming through coral reef. Roatan Island, Honduras. Caribbean Sea
Yellowtail snapper (Ocyurus chrysurus) hooked on fishing line. Yellowtail snapper fishery, Utila Island, Honduras. Caribbean Sea
Low angle view of Jaguar (Panthera onca) patrolling its territory at night, La Papalota, Nayarit, Mexico. Camera trap image
Silhouette of Axolotl (Ambystoma mexicanum) swimming in a woodland stream surrounded by towering trees, Sierra de Manantlan, Mexico
Arctic tern (Sterna paradisaea) in flight over nesting area in meadows near coast, Reykjanes Peninsula, Iceland. July
Male Lemon shark (Negaprion brevirostris) gliding just below the surface, Grand Bahama, Caribbean Sea
Palm-nut vulture (Gypohierax angolensis) in flight, Allahein river, The Gambia
Small flock of migrating Black-tailed godwits (Limosa limosa) in flight against a bright blue sky, River Coquet Estuary, Northumberland, UK. September
Wintery forest of Silver Birches (Betula pendula) photographed from below with a wide angle lens. Białowieła National Park, Poland. January
Ventral view of Swallowtail butterfly (Papilio machaon) in flight. The Netherlands. July
Steller sealion (Eumetopias jubatus) swimming over seabed, Vancouver Island, British Columbia, Canada, Pacific Ocean
Male Great frigatebird (Fregata minor) in breeding condition with semi-inflated throat pouch, Christmas Island, Australia
Santa Fe land iguana (Conolophus pallidus) with Giant prickly-pear cactus (Opuntia echios), Santa Fe Island, Galapagos, South America
Venomous Yellow-lipped sea krait (Laticauda colubrina) heading up to the surface to breathe, Philippines
Female Red fox (Vulpes vulpes) foraging inside a rotting tree trunk, Verteskozma, Hungary. Hungary Nature Photographer of the Year Competition 2021 Highly Commended - Mammals Behaviour category
Spider crab (Hyas sp. ) scaling the stipe of Cuvie kelp (Laminaria hyperborea)Spider crab (Hyas sp.) scaling the stipe of Cuvie kelp (Laminaria hyperborea) up into the canopy of the kelp forest. Loch Carron, Highlands, Scotland, UK
European beech forest (Fagus sylvatica) in autumn, view from below in the mist, Ilirska Bistrica, Green Karst, Slovenia, October
Northern kelp crab (Pugettia producta) clinging to a fallen tree branch off Vancouver Island, British Columbia, Canada
California redwoods (Sequoia sempervirens) planted in 1939, Beech Forest, Victoria, Australia
Common carder bumblebee (Bombus pascuorum) nectaring on a Dandelion (Taraxacum officinale) flowerhead among many Common daisies (Bellis perennis)
Ragged robin (Lychnis flos-cuculi) flowering in wet flower-rich meadow, Kielder Water and Forest Park, Northumberland, UK, June 2020
Long-nosed Potoroo (Potorous tridactylus) eating fungi. Captive, Conservation Ecology Centre, Victoria, Australia. June 2017
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Honey fungus (Armillaria mellea) clumps in deciduous woodland, Buckholt Wood NNR, Gloucestershire, UK, October
Red stingray (Hemitrygon akajei) with Banded houndshark (Triakis scyllium) group and smaller fish in background. Chiba Prefecture, Honshu, Japan. September
Red stingray (Hemitrygon akajei), four in a row, viewed from below. Chiba Prefecture, Honshu, Japan. October
Lemon shark (Negaprion brevirostris) hunting over Turtlegrass (Thalassia testudinum) seagrass bed, view from below. Florida Keys, Florida, USA
Green surf anemones (Anthopleura xanthogrammica) grow in shallow water below the temperate rain forest. Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada. North East Pacific Ocean
Coastal agave (Agave shawii), two flower spikes against sky, view from below. Near Bahia de los Angeles. Baja California, Mexico
A lesser horseshoe bat (Rhinolophus hipposideros) flying over grassland habitat at night. Devon, UK
A large female Grey seal (Halichoerus grypus) probably heavily pregnant, resting/bottling at the surface in shallow water, head above water, Farne Islands, Northumberland, UK
Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February
Monarch butterflies (Danaus plexippus) flying during a warm morning. When the overwintering period draws to an end large numbers become active. Michoacan, Mexico, February
Red squirrel (Sciurus vulgaris) feeding on Maple (Acer sp) seeds in tree. Jyvaskyla, Central Finland. November
Siberian jay (Perisoreus infaustus) flying in coniferous forest. Kuusamo, Northern Ostrobothnia, Finland. February
Mahogany treefrog (Tlalocohyla loquax) clasping onto leaf stem, view from below. Las Guacamayas Biological Station, Laguna del Tigre National Park, El Peten, Guatemala
King penguins (Aptenodytes patagonicus) on the beach at Salisbury Plain, with Antarctic Tern (Sterna vittata) hovering. South Georgia, South Atlantic
Fleecy milk-cap fungus (Lactifluus / Lactarius vellereus) on the forest floor in beech woodland in autumn, France, November
Scalesia forest canopy (Scalesia pendunculata), Los Gemelos, Santa Cruz Highlands, Galapagos Islands
Oceanic whitetip shark (Carcharhinus longimanus) is framed by the surface. Elphinstone reef, Marsa Alam, Egypt. Red Sea
Summer snowflake (Leucojum aestivum), restricted to the Thames Valley. Withymead, near Goring, Oxfordshire, England, UK, April
Herb paris (Paris quadrifolia) in Lower Woods, Gloucestershire, UK. April
Common snowdrops (Galanthis nivalis) growing in the Rococo Gardens, Painswick, Gloucestershire, Engalnd, UK. February
Stormy sky over Klein Schietveld, Brasschaat, Belgium. March 2019
American oak (Quercus rubra) trees in avenue, sun shining through. Peerdsbos, Brasschaat, Belgium. November 2013
Black and white ruffed lemur (Varecia variegata) hanging from tree, Madagascar, critically endangered
View upward to canopy of a Beech woodland (Fagus sylvatica) in autumn. Plitvice Lakes National Park, Croatia. October
View upwards to canopy of a Beech woodland (Fagus sylvatica) in autumn. Plitvice Lakes National Park, Croatia. October
