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Deciduous forest with sunlight shining through trees, Monmouthshire, Wales, UK. September
Leopard houndshark (Triakis semifasciata) group swimming amongst rays of light. San Diego, California, USA
Lemon shark (Negaprion brevirostris) hunting over Turtlegrass (Thalassia testudinum) seagrass bed, view from below. Florida Keys, Florida, USA
Langur monkey (Semnopithecus) climbing tree in misty forest at dawn, India, November
Oceanic whitetip shark (Carcharhinus longimanus) swims in open waters, close to the surface with sun beams. Elphinstone reef, Marsa Alam, Egypt. Red Sea
Young female Grey seal (Halichoerus grypus) beneath sunburst. Farne Islands, Northumberland, UK
Chacma baboon (Papio ursinus) sitting in rainforest in morning light. Jao Reserve, Botswana
Sunbeams play through the roots of red mangrove trees (Rhizophora sp.), encrusted with marine life and home to schoolmaster snappers (Lutjanus apodus)
RF - American crocodile (Crocodylus acutus) swimming through sunrays, above a bed of seagrass ( Thalassia testudinum). Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
RF - Wandering albatross (Diomedea exulans) flying over the rugged coast of South Georgia. South Atlantic. (digitally stitched image)
Early morning view, looking southeast from Gordon Channel. Gordon Channel, Queen Charlotte Strait, British Columbia, Canada. September
Sperm whale (Physeter macrocephalus) mother and calf, Sri Lanka. March
Meadow of Neptune seagrass (Posidonia oceanica), Chersonissos, Heraklion, Crete, Greece
Neptune seagrass (Posidonia oceanica) meadow, Samaria National Park, Chania, Crete, Greece
Autumn crocus or meadow saffron (Colchicum autumnalis) in Lower Woods, Gloucestershire, England, UK, September
Rays of sunlight through mixed Beech (Fagus sp) / Oak (Quercus sp) forest after heavy downpour of hail, Peerdsbos, Brasschaat, Belgium, June 2014
Sun rays filter through row of Oak (Quercus robur) trees. Peerdsbos, Brasschaat, Belgium. November
American oak (Quercus rubra) trees in avenue, sun shining through. Peerdsbos, Brasschaat, Belgium. November 2013
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Sunbeams shining through Bamboo (Bambusidae) leaves. Shunan Zhuhai National Park, Sichuan Province, China
Early morning shafts of light through deciduous forest. Bandhavgarh National Park, India
Whale shark (Rhincodon typus) front view portrait, Cenderawasih Bay, West Papua. Indonesia
Rays of light in a fen on a misty morning, Oisterwijkse Bossen en Vennen Reserve, Oisterwijk, The Netherlands, January
The Quiraing, Trotternish Peninsula, Isle of Skye, Inner Hebrides, Scotland, UK. January 2015
Lagoa Misteriosa at the bottom of a sink hole in limestone region, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil November 2016. Photographed for The Freshwater Project
Brydes whale (Balaenoptera edeni) tail, Trincomalee, Eastern Province, Sri Lanka, Bay of Bengal, Indian Ocean
RF- (Pinus sylvestris) at sunrise, Klein Schietveld, Brasschaat, Belgium
Scots pine (Pinus sylvestris) forest with sun filtering through mist, Alvie Estate, Kincraig, Cairngorms National Park, Scotland, UK, June
Light streaming through window on sand covered house in Kolmanskop Ghost Town, an old diamond-mining town where shifting sand dunes have encroached abandoned houses, Namib Desert Namibia, October 2013
Barrel Jellyfish (Rhizostoma pulmo) against crepuscular light rays. Sark, British Channel Islands, July
RF - Silky shark (Carcharhinus falciformis) swimming beneath the surface of the ocean, with evening sun burst. Gardens of the Queen National Park, Cuba
Grey seal (Halichoerus grypus) young seal silhouetted against the sun as it dives after breathing, Lundy Island, Devon, UK, Bristol Channel, August
Broadclub cuttlefish (Sepia latimanus) Komodo National Park, Indonesia
Rays of sunlight penetrating woodland, Rhinefield, The New Forest, Hampshire, UK. November 2012
Three White rhinoceroses (Ceratotherium simum), Solio Game Reserve, Laikipia, Kenya. September