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Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Cavalier King Charles Spaniel, bitch with black-and-tan coat
Kelp forest (Laminaria hyperborea) in the clear seas of Scotland, UK, September
Tibetan antelope (Pantholops hodgsonii) male, Kekexili, Qinghai, Tibetan Plateau, China, November
Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m
American paddlefish (Polyodon spathula), an introduced species native to the Mississippi River Basin, USA, swimming over lake bed, private lake, Moscow region, Russia
Long-tailed potoo (Nyctibius aethereus) camouflaged on roosting perch in lowland rainforest, Manu Biosphere Reserve, Peru
Sandstone long-necked turtle (Chelodina burrungandjii) actively foraging at night, Pentecost River, Western Australia. April
Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
White rhinoceros (Ceratotherium simum) with a long horn, Marakele National Park, South Africa
Resplendent quetzal (Pharomachrus mocinno) male perched on branch. Costa Rica
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Common / Eurasian cranes (Grus grus) taking flight for roasting site, at sunset, silhouetted. Lake Hornborga, Sweden, April
Sword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devil sSword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devils Trumpet Flower (Datura sp.). Yanacocha montane cloud forest, Ecuador
Reeftop pipefish (Corythoichthys haematopterus) swimming over hard coral (Diploastrea heliopora), Laamu Atoll, Maldives, Indian Ocean
White rhinoceros (Ceratotherium simum) with unusually shaped horn, Solio Game Reserve, Laikipia, Kenya. September
Gannet (Morus bassanus) chick with fluffy with mouth open. Shetland Islands, Scotland, UK, August
Greater yellowlegs (Tringa melanoleuca) standing on rock. Acadia National Park, Maine, USA. September
RF- Walrus (Odobenus rosmaris) close up head portrait, Svalbard, Norway
Great Grey owl (Strix nebulosa) taking off from the ground in snow, Finland, April. Bookplate from Danny Greens The Long Journey North