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Glass frog (Rulyrana spiculata) ventral / underside view, Cosnipata Valley, Peru
Indian moon / Indian luna Moth (Actias selene) close up of defensive spines on caterpillar. Captive
Common Swallowtail Butterfly (Papilio machaon) resting on Vipers Bugloss / Blueweed (Echium vulgare) in alpine meadow. Nordtirol, Tirol, Austrian Alps, Austria, 1700 metres altitude, July
Silhouette of spider at sunset, Berwickshire, Scotland, August
Elephant hawk moth in flight {Deilephila elepenor} UK
Bigfin squid (Sepioteuthis lessoniana) hovering in mid water at night. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Golden damselfish (Amblyglyphidodon aureus) hiding amongst the branches of red whip coral (Ctenocella sp.) Misool, Raja Ampat, West Papua, Indonesia
Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Chocolate-dip chromis (Chromis dimidiata) male over out of focus hard coral (Porites nodifera). Sha ab Claudia, Fury Shoal, Egypt. Red Sea
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon, on coral reef under stormy seas. St Johns Reef. Egypt. Red Sea
Pink shrimp (Pandalus montagui), Loch Carron, Ross and Cromarty, Scotland, UK, April
Secretary blenny (Acanthemblemaria maria) making a home in a deserved worm tube of a great star coral (Montastraea cavernosa), East End, Grand Cayman, Cayman Islands, British West Indies
Two male Common cuttlefish (Sepia officinalis) compete for a female (on right) during courtship, Torbay, Devon, UK, May. 2020VISION Exhibition. 2020VISION Book Plate
A Mountainous star coral (Montastraea faveolata) spawning at night, with gamete bundle being released from a coral polyp, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea
RF - Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Studio image of multiple Nudibranch species. Gulen, Norway. North East Atlantic Ocean. Digital composite
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
Common blue butterfly (Polyommatus icarus), resting on flowering heather, Arne RSPB reserve, Dorset, England, UK September
Pink anemonefish (Amphiprion perideraion) in a purple Magnificent sea anemone (Heteractis magnifica). Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Red boring sponge (Cliona delitrix) three excurrent apertures covered in Sponge zoanthids (Parazoanthus parasiticus) East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Bullseye / Io moth (Automeris io) showing eye spot markings on wings during deimatic display to deter predators, originating from North and Central America, sequence 2 of 2. Captive
Yellow cluster anemone (Parazoanthus axinellae) surrounded by Encrusting sponge (Spirastrella cunctatrix) San Pietro Island, Sardinia, Italy, Mediterranean, July
Starfish close up detail, Blairgowrie, Mornington Penisular, Victoria, Australia. Port Philip Bay, Bass Strait
Garden snail (Helix aspersa) baby on Marigold flower, UK
Close-up of the polyps out of a coral tree (Tubastraea coccinea) at night, Baja California peninsula, Mexico. Sea of Cortez
Orange cup coral (Tubastrea coccinea) Malpelo Island National Park, UNESCO World Heritage Site, Colombia, East Pacific Ocean
Red Admiral Butterfly (Vanessa atalanta) on Michaelmas daisy flowers. Dorset, UK, October 2012
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
A tiny amphipod (Iphimedia obesa) living on Dead mans fingers soft coral (Alcyonium digitatum) Loch Carron, Ross and Cromarty, Scotland, UK, April
Female Downy Emerald dragonfly (Cordulia aenia) at rest on flowering Gorse (Ulex europaeus) Higher Hyde Heath, Dorset, UK. April
Brimstone butterfly (Gonepteryx rhamni) Male at rest on Primrose flower, West Sussex, UK April
Emperor angelfish (Pomacanthus imperator) swimming past soft corals (Dendronephthya sp]), Red Sea, Egypt. January
Flatworm (Maiazoon orsaki) Raja Ampat, Irian Jaya, West Papua, Indonesia, Pacific Ocean
Two-tailed Pasha butterfly (Charaxes jasius) expanding wings after emerging, Podere Montecucco. Orvieot, Umbria, Italy August. Sequence 9 of 9
Light Bulb Sea Squirt (Clavelina lepadiformis). Channel Islands, UK, June
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
Brain coral (Diploria strigosa) photographed under blue light to excite the fluorescence, Little Cayman, Cayman Islands, British West Indies, Caribbean Sea
Blue sea star (Linckia laevigata) on coral reef, Fiji, South Pacific
RF - Long-spined sea urchin (Astropyga radiata). Lembeh Strait, North Sulawesi, Indonesia
Gorgonian coral (Acanthogorgia sp. ) Similan Islands, Andaman Sea, ThailandGorgonian coral (Acanthogorgia sp.) Similan Islands, Andaman Sea, Thailand
Brown-lipped snail (Cepaea nemoralis) on fungus, reproduced in a bokeh affect, Sheffield, England, UK. November. Commended in the Natures Studio Category of GDT competition 2014