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Long-jawed orbweaver (Metellina segmentata) in web at sunset. Dunwich Heath, Suffolk, UK, September
Small tortoishell butterfly (Aglais urticae) on a stained glass window of church on a sunny day Sheffield, England, UK, January
Blue tit (Cyanistes caeruleus) bringing Crab spider prey (Misumena vatia) to feed chicks, Monmouthshire, Wales, UK. May
Portrait of Blacktip grouper (Epinephelus fasciatus) resting in Barrel sponge (Xestospongia testudinaria) Mabul Island, Sabah, Borneo, Malaysia. Sulawesi Sea
Queen conch (Strombus gigas) extending its proboscis and stalked eyes at it moves across the sand by coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Lions mane jellyfish (Cyanea capillata) with feeding tentacles spread swimming in the sun in shallow water. Gulen, Norway. North East Atlantic Ocean
Shallow coral reef flat with corals (Pocillopora damicornis, Acropora humilis and Pocillopora verrucosa) in evening light. Gordon Reef, Strait of Tiran, Sinai, Egypt. Gulf of Aqaba, Red Sea
Hooded nudibranchs (Melibe leonina) filter feeding beneath bull kelp. Hussar Point, Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
Spiny spider crab (Achaeus spinosus) reaching up and scratching its head, while climbing on purple soft coral bush (Dendronephthya sp.) Sinai, Egypt. Red Sea
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
Oman anemonefish / clownfish (Amphiprion omanensis) in a a Bulb-tentacle sea anemone (Entacmaea quadricolor), coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Marbled White butterfly (Melanargia galathea) resting on grass, Dunsdon Nature Reserve, Devon, UK. July
Common hawker (Aeshna juncea) Brackish Moss NNR, County Armagh, Northern Ireland
Banded demoiselle (Calopteryx splendens) male close up detail of head and thorax, Hertfordshire, England, UK, June. Captive. Focus stacked
RF - Marbled white butterfly (Melanargia galathea) on knapweed, with soft focus leaves, Aosta Valley, Gran Paradiso National Park, Italy
Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August
Leaf cutter ants (Atta sp) carrying pieces of fern, reflected in water, Costa Rica
Stag beetle (Lucanus cervus) male silhouetted against leaf, controlled conditions
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
Morpho (Morpho peleides) eggs, Green Hills Butterfly Ranch, Cayo District, Belize
Blue iridescent scales of a Nymphalid butterfly (Epiphile orea) magnified 11x, deceased insect found in Piedade, Sao Paulo, Brazil. South-east Atlantic Forest
Small tortoiseshell butterfly (Aglais urticae) resting on old painted door. Dorset, UK, August
Buff-tailed bumblebee (Bombus terrestris) flying to Oriental poppy (Papaver orientale) Monmouthshire, Wales, UK. July
Luminescent jellyfish / Mauve stinger (Pelagia noctiluca) in open water, Gozo Island, Malta. Mediterranean Sea
Green praying mantis (Majangella moultoni) camouflaged on mossy tree trunk. Danum Valley, Sabah, Borneo
Merveille-du-Jour moth (Dichonia aprilina), camouflaged on lichen. The National Forest, Leicestershire, UK. September
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
Detail of rhinopores and the colour pattern of a toxic Nudibranch (Goniobranchus coi) on a coral reef. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink
Red Sea anemonefish (Amphiprion bicinctus) in their home, a Sea anemone (Heteractis magnifica), on a coral reef. Big Brother island, Red Sea. Egypt
Hercules moth (Coscinocera hercules) recently emerged in montane rinforest. Ambua Lodge, Tari, Hela Province, Papua New Guinea. June
Squid (Abraliopsis atlantica), with photophores in 6 straight longitudinal rows on mantle visible. They are used to produce bioluminescencent light
Juvenile Squid (Onychoteuthis sp. ) deep sea species from Atlantic Ocean off Cape VerdeJuvenile Squid (Onychoteuthis sp.) deep sea species from Atlantic Ocean off Cape Verde. | Captive
Squid (Histioteuthis sp. ) deep sea species from Atlantic Ocean off Cape Verde. CaptiveSquid (Histioteuthis sp.) deep sea species from Atlantic Ocean off Cape Verde. Captive
Giant deep-sea isopod (Bathynomus giganteus) Picture was taken in cooperation with the Zoological Museum University of Hamburg
(Cione limacina) sea angel, a pelagic pteropod mollusc from the mid-Atlantic, deep sea Atlantic ocean
Peacock butterflies (Inachis io) feeding on corn marigolds (Glebionis segetum) on agricultural headland. England, UK
Fringed Tulip Dallas (Pink) and Tulipa Swan Wings (White) and Tulipa Valery Gergiev (red) growing in garden border. England, UK
Thimble jellyfish (Linuche unguiculata) aggregation, My Paradise Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Three False clownfish (Amphiprion ocellaris) in Sea anemone (Heteractis magnifica) Lighthouse Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Tropical shield mantis (Choeradodis rhombicollis) in defensive position, on flower, Chinambi, Carchi, Ecuador
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise with bivalve shell
Chapuline grasshopper (Sphenarium mexicanum) portrait. Los Tuxtlas rainforest, Mexico, . July
Mount Lewis spiny crayfish (Euastacus fleckeri) in defensive posture with claw raised, at edge of mountain stream, Mount. Lewis, Wet Tropics World Heritage area, Queensland, Australia. Endangered
Caterpillar, probably Cholla moth caterpillar, (Euscirrhopterus cosyra) on cactus, Texas, USA. May
Compass jellyfish (Chrysaora hysoscella) feeding on plankton near surface, Falmouth, English Channel, Cornwall, UK. July
Two Asian weaver ants (Oecophylla smaragdina), queens founding a new nest, tending to eggs, West Bengal, India