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Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Whitetip reef sharks (Triaenodon obesus) pack silhouetted following scent trail in water column, Cocos Island, Costa Rica, Pacific Ocean
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
Red Kite (Milvus milvus) diving towards prey whilst in flight, Wales, UK, March
Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive
Cheetah (Acinonyx jubatus) running at full speed chasing Springbok prey, Kalahari Desert, Southern Africa
Golden Eagle (Aquila chrysaetos) swooping with folded wings and extended tallons. Canyon del Ebro y Rudron, Castilla y Leon, Spain. Digital composite
Grizzly bear (Ursus arctos horribilis) catching salmon in Brooks river, Katmai National Park, Alaska, USA, July
Osprey (Pandion haliaetus) with two Alewife (Alosa pseudoharengus) just caught in the Atlantic Ocean. Acadia National Park, Maine, USA. June