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Snowy owl (Bubo scandiaca) female, with lights behind, Canada, February
Killer Whales / Orcas (Orcinus orca)Killer whales / Orcas (Orcinus orca) large pod including calf traveling together while foraging on large schools of Herring (Clupea harengus) in the cold waters of northern Norway, January
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Grey seal (Halichoerus grypus) swimming towards camera, Orkney, Scotland, UK, August
European roller (Coracias garrulus) in flight, Pusztaszer, Hungary, May
Head portrait of grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
Head portrait of black Merens stallion with long mane running in pasture. Northern France, Europe. February
Fin whale (Balaenoptera physalus) near surface. Pico Island, Azores, Portugal. May
Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Fieldfare (Turdus pilaris) in Rowan tree with berries in winter, Jyvaskyla, Central Finland, January
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Scots pine (Pinus sylvestris) with Northern lights / Aurora borealis lighting up the night sky in background, Monadhliath Mountains, Cairngorms National Park, Scotland, UK, October 2015
Cockapoo, Cocker spaniel cross Poodle puppy sitting
Yellow Labrador Retriever puppy, 8 weeks old, sitting with raised paw
Blue British Shorthair kitten sitting
Tabby kitten, 6 weeks, standing
Proboscis monkey (Nasalis larvatus) Riverine forest, Kinabatangan River, Sukau, Sabah, Borneo
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Warthog (Phacochoerus aethiopicus) female with piglets, Okavango Delta, Botswana
Male Lion (Panthera leo) portrait, close-up of face, captive, occurs in Africa
Tabby cats, Picasso and Smudge, age 3 months, together. Smudge with his paws up on Picasso
Red squirrel (Sciurus vulgaris) stood on log in snow, Cairngorms National Park, Scotland, UK. December
Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA
Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August
Minke whale (Baelanoptera acutorostrata) breaching, Bay of Fundy, New Brunswick, Canada, July
Common vampire bats (Desmodus rotundus) roosting in cave, Costa Rica
Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean
North American Bobcat (Lynx rufus) stalking along Madison River. Yellowstone National Park, Wyoming, USA. January
Greater roadrunner (Geococcyx californianus), adult drinking, Rio Grande Valley, South Texas, Texas, USA. June
European flamingo (Phoenicopterus roseus) bathing Camargue, France, May
Black and white Jack-a-poo, Jack Russell cross Poodle, pup, 8 weeks old, jumping up
Grey seal (Halichoerus grypus) swimming over kelp forest, Shetland, Scotland, UK, August
Bighorn sheep (Ovis canadensis) close up rams head shot, Yellowstone National Park, USA, February
Snowy owl (Bubo scandiaca) female face close up, Canada February
Red deer (Cervus elaphus) stag in autumn woodlands, Bradgate Park, Leicestershire, UK, October
Grey seal (Halichoerus grypus) split level of young seal investigating the camera with its nose and whiskers, close to shore. Lundy Island, Devon, UK, Bristol Channel. August
Swallowtail butterfly (Papilio machaon) in garden, Norfolk, England, UK, June
Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016
Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Lightning storm over Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May 2012
Arabian Oryx (Oryx leucoryx) at sunset. Previously extinct in the wild, their conservation status is now stable thanks to conservation efforts.This male is part of a reintroduction program in Dubai
Emerald tree Boa (Corallus batesii) close up portrait of juvenile whilst hanging on a branch. Tambopata, Madre de Dios, Peru
Two Ladys slipper orchids (Cypripedium calceolus) in flower in spring in woodland near Kuressaare, Saaremaa, Estonia
Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August