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Fallow deer (Dama Dama) stags in velvet standing in a shady woodland spot, Knepp Estate, Sussex, UK, June
Herd of domestic Water buffalo (Bubalus arnee) in dust at dusk, Lake Kerkini area, Greece, July
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Northern Serenget National Park, Tanzania (early September)
Angolan giraffes (Giraffa giraffa angolensis) approaching a scarce waterhole. Etosha National Park, Etosha, Namibia
Konik horses (Equus ferus caballus) interacting at sunset, floodplain of the river Rijn, Meinerswijk near Arnhem, the Netherlands. May
Blue sheep (Ovis nahoor) Angsai Nature Reserve (Valley of the Cats), Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China
Kiang or Tibetan Wild ass (Equus kiang) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
A herd of wild konik horses running, with refection in water, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Yak (Bos grunniens) herd on plain beside river, mountains in background. Near Litang, Kham, Garze Tibetan Autonomous Prefecture, Sichuan, China. October 2016
European bison (Bison bonasus) herd in grassland. Eriksberg Wildlife and Nature Park, Blekinge, Sweden. June. Captive
Wild buffalos (Bubalus arnee) on beach in Pui O, Lantau Island, Hong Kong, China
Lusitano horses, five young stallions playing between trees in pasture, silhouetted at dawn. Portugal. April
RF - Dulmen pony, wild mares and foals running together at roundup. Dulmen, North Rhine-Westphalia, Germany. (This image may be licensed either as rights managed or royalty free.)
African elephant (Loxodonta africana) herd drinking at a waterhole, Mashatu Game Reserve, Botswana. June
African buffalo (Syncerus caffer) herd showing defensive behaviour as an African lioness (Panthera leo) walks by, Mana Pools National Park, Zimbabwe. September
Horses running in grassland. Bashang Grassland, near Zhangjiakou, Hebei Province, Inner Mongolia, China. July
RF-Horses running through Bashang Grassland. Near Zhangjiakou, Hebei Province, Inner Mongolia, China. July. (This image may be licensed either as rights managed or royalty free)
RF - Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Serengeti National Park, Tanzania. September
Nile crocodile (Crocodylus niloticus) waiting to attack White-bearded wildebeest (Connochaetes taurinus albojubatus) as they cross the Mara River
RF - Impala (Aepyceros melampus) at water with reflections, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Springbok (Antidorcas marsupialis) herd on the move, Kgalagadi transfrontier park, South Africa
African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
African lion, (Panthera leo) confronted by a herd of African buffalo / Cape buffalo (Syncerus caffer), Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Fallow deer (Dama dama) herd at dawn, Holkham, Norfolk, England, UK, October
Red deer (Cervus elaphus) herd of female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) female does and young crossing water, Jura, Scotland, UK, September
Himalayan ibex (Capra sibirica) during a snowfall, coming down the mountain at dusk to sleep at lower altitude, around Kibber village, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains
Aerial view of African elephants (Loxodonta africana) looking for the shade under trees on a very hot day, Northern Botswana
Plain / Burchells zebras in dust (Equus burchelli) Nxai Pan National Park, Botswana
Zebra at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
European bison / Wisent (Bison bonasus) herd released into the Tarcu mountains nature reserve, Natura 2000 area, Southern Carpathians, Romania. May 2014
RF- Caribou herd (Rangifer tarandus) grazing on tundra. Kobuk Valley National Park, Alaska, USA
Wild yak herd (Bos mutus) Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
BlackbuckA(Antilope cervicapra), herd with males and females, Velavadar National ParkBlackbuck(Antilope cervicapra), herd with males and females, Velavadar National Park, Gujarat, India
Greater blue sheep (Pseudois nayaur), herd on mountain slope, important prey for snow leopards (Panthera uncia), Himalaya, Hemis National Park, Ladakh, India
Silhouette of herd of female Red deer (Cervus elaphus) on ridge at dawn, RSPB Forsinard Flows, Flow country, Caithness, Highland, Scotland, UK, June 2011
Herd of Red deer (Cervus elaphus) stags in clearing in snow covered pine forest, Cairngorms National Park, Scotland, UK, December
Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya
Herd of Impala (Aepyceros melampus) silhouetted at sunset, Ngorongoro Conservation Area, Tanzania
Fallow deer (Dama dama) one deer stops and looks whilst the herd runs on, with sunlit leaves out of focus, Vastmanland, Sweden
Charolais cattle on eroded river bank, Monmouthshire, Wales, UK, August
Herd of Camargue bulls, Camargue, France, May
A group of Pure Arab, Shagya Arab and East Bulgarian fillies running in snow, Kabiuk National Stud, Shumen, Bulgaria
Springbok (Antidorcas marsupialis) herd, Kgalagadi Transfrontier Park, South Africa
Bison (Bison bison) herd walking in line in snow, Yellowstone National Park, Wyoming, USA, February
