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Brown Hare (Lepus europaeus) portrait. Derbyshire, UK, March
European river otter (Lutra lutra) resting amongst the seaweed, Isle of Mull, Inner Hebrides, Scotland, UK, December
European river otter (Lutra lutra) swimming in the sea, Isle of Mull, Inner Hebrides, Scotland, UK, December
Northern gannet (Morus bassanus) portrait, Saltee Islands, Ireland, May 2008
Portrait of a Hawk Owl (Surnia ulula). Uto, Finland, March
Red fox (Vulpes vulpes) alert cub portrait, Derbyshire, UK, June. Non-ex
Grass snake (Natrix natrix) swimming amongst duck weed, Oxfordshire, UK
Red fox (Vulpes vulpes) portrait, England
Tawny owl (Strix aluco) in amongst hawkbit flowers, Leontodon, UK, September. Captive
RF- Northern Goshawk (Accipiter gentilis) male close-up, Scotland
Red legged partridge (Alectoris rufa) in corn field, Norfolk, UK, July. Non-ex
Gannet (Morus bassanus) portrait, Shetland Islands, Scotland, UK
Large / Pot Bellied Seahorse (Hippocampus abdominalis) face portrait. Blairgowrie Marina, Mornington Peninsular, Victoria, Australia, March
Male Azure damselfly (Coenagrion puella) resting on a leaf, Vallee de la Seille (Seille Valley), Lorraine, France, June
Beluga whale (Delphinapterus leucas) swimming under ice, Arctic circle Dive Center, White Sea, Karelia, northern Russia captive
Amazon river dolphin (Inia geoffrensis), Rio Negro, Amazonia, Brazil
Portrait of Southern Elephant Seal (Mirounga leonina), the largest species of pinniped. South-Georgia, Antarctica, January
Head of Beluga / White Whale (Delphinapterus leucas) in profile. Canada, summer. Captive
Two Konik Wild Horse (Equus ferus caballus) interacting. The Netherlands, November
Beluga / White whale (Delphinareptus leucas) surfacing, Captive, Vancouver Aquarium, Canada
Meerkat looking into lens {Suricata suricatta} Tswalu Kalahari Reserve, South Africa
Head portrait of Satiny Parrot snake (Leptophis depressirostris) swallowing frog, Hungary, Europe
African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
Lammergeier (Gypaetus barbatus) close up portrait, Spain, November
Northern gannets (Morus / Sula bassanus) portrait of individual with a courting pair in foreground, UK, Bass Rock, Scotland
Golden Eagle (Aquila chrysaetos) portrait in snow, Utajarvi, Finland February
Polar Bear (Ursus maritimus) head portrait with blue tongue sticking out, Svalbard, Norway
Male Marsh pride lion (Panthera leo) on a moonless night, Masai Mara, Kenya. Taken with infra red camera, September
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Close-up of Brown bear (Ursus arctos) in lake, Kamchatka, Far east Russia, August
Highland bull (Bos taurus) face close up in freezing fog, Berwickshire Scotland, December
Clouded leopard (Neofelis nebulosa) Guwahati, Assam, India, captive
Southern plains grey langur ( Semnopithecus entellus / Presbytis entellus) close up face profile, Sawai Modhopur, Rajasthan, India
Boxer looking over closed door, standing on hind legs
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Red river hog (Potamochoerus porcus) close up of head, captive
Red Grouse (Lagopus lagopus scoticus) in Heather. Peak District, UK, August
Red fox (Vulpes vulpes) head portrait, The Netherlands, March
Wild cat (Felis silvestris) in snowstorm captive, Germany
German Hound male, 6 months
Black swan (Cygnus atratus) captive
Comtois Horse (Equus caballus). Camargue reserve, Rhone, France, September
Southern elephant seal (Mirounga leonina) close up of sleeping individual, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Grey seal (Halichoerus grypus) adult female among breaking waves, Lincolnshire, UK Non-ex
Yellow Labrador Retriever puppy, 7 weeks
A portrait of a Leafy Seadragon (Phycodurus eques). Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Border Collie puppy portrait