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Primrose (Primula vulgaris) in Lower Woods, Gloucestershire, England, UK, March
Clarkes pool meadow, an unimproved lowland hay meadow managed by the Gloucestershire Wildlife Trust, dotted with Green-winged orchids (Anacamptis morio) in early spring
Common cotton grass (Eriophorum angustifolium) Klein Schietveld, Brasschaat, Belgium, May
Red deer (Cervus elaphus) stag in snowy pine forest. Cairngorms National Park, Highlands, Scotland, UK, March. Highly commended in the Wild Woods Category of the BWPA Awards 2019
Catalina Desert Iguana (Dipsosaurus catalinensis), Catalina (Santa Catalina) Island, Loreto Bay National Park, Gulf of California (Sea of Cortez), Baja California, Mexico, May
Red-and-green macaw (Ara chloropterus) flock feeding at wall of clay lick, flying and perched in trees. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Emergent Menggaris Tree / Tualang (Koompassia excelsa) protruding from mist and low cloud hanging over lowland rainforest. Danum Valley, Sabah, Borneo
Juvenile Reticulated python (Python reticulatus) resting coiled round sapling by a stream. Danum Valley, Sabah, Borneo, Malaysia
Icebergs and autumn tundra near Rode O (Red Island) in Rode Fjord (Red Fjord), Scoresby Sund, Greenland, August
Polar bear (Ursus maritimus) walking across ice, snow covered mountain in background. Svalbard, Norway, April 2018
RF - Guanaco (Lama guanicoe) standing in front of mountain towers of Paine. Torres del Paine National Park, Patagonia, Chile. December 2018
Guanaco (Lama guanicoe) standing, towers of Torres del Paine National Park in background. Patagonia, Chile. December 2018
Namaqua dwarf adder, (Bitis schneideri), Little Namaqualand, South Africa, February. Non-ex
Johnston's three-horned chameleon, (Trioceros johnstoni), male on tree branch, Kahuzi-Biega NP, Democratic Republic of Congo, November. Non-ex
Quiver tree forest (Aloe dichotoma) at night with stars and the Milky Way, Keetmanshoop, Namibia
Grand-flowered horned poppies (Glaucium grandiflorum) in southern Turkey, June
Black Forest tree trunks in mist, Baden-Wurttemberg, Germany. May
Aerial image of Dehesa forest, Salamanca Region, Castilla y Leon, Spain, May 2011
Roe Deer (Capreolus capreolus) hiding behind a tree. Black Forest, Germany, May
Short-toed Eagle (circaetus gallicus) flying amongst Stone Pines (Pinus pinea). Coto Donyana, Andalusia, Spain. 2007
Caatinga tree. Note thorns. Drought resistant species, storing water in swollen trunk. Bahia State, NE Brazil. South America. Threatened Habitat
Crested tit (Lophophanes cristatus) perched above pine forest with a series of small lochans in background, Cairngorms National Park, Scotland, UK, December 2014
RF - European ground squirrels / Sousliks (Spermophilus citellus)greeting, Gerasdorf, Austria. April
Autumn colours of Poplar (Populus sp) Ordesa y Monte Perdido National Park, Spain. October
Red fox (Vulpes vulpes) in autumnal colours, captive
Fallow deer (Dama dama) buck bellowing in autumn vegetation, Bradgate Park, Leicestershire, UK, October
Oak woodland with mosses and running water, near Dolgellau, Snowdonia, North Wales, UK October
Alpine ibex (Capra ibex) on a mountain side in autumn with European larch tree (Larix decidua) trees in the back the Alps, Gran Paradiso National Park, Italy. November
Quinling Golden snub nosed monkey (Rhinopitecus roxellana qinlingensis) family group foraging along a small creek in a gullly. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
Scattered Scots pine trees (Pinus sylvestris) beside the Dorback Burn, Dorback Estate, Cairngorms National Park, Scotland, UK, September 2011
Red Fox (Vulpes vulpes) behind a tree and autumn leaves. Black Forest, Germany, November
Autumns colours in Carpathian beech forest. Bieszczady, Carpathian Mountains, Poland, November
Woodland stream flowing through Wilmersham Wood, Exmoor National Park, Somerset, UK. November 2010
Group of Wild Boar (Sus Scrofa) trotting through forest of Rambouillet, near Paris, France. Autumn
Sooty albatross (Phoebetria fusca) nesting amongst Blechnum palmiforme tree ferns. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos
Galapagos hawk (Buteo galapagoensis) perched over sulphurous slope, Galapagos, Ecuador. Vulnerable species
Lowland blotched blue-tongue (Tiliqua nigrolutea) on a coastal bluff in the Bay of Fires region of north-eastern Tasmania, Australia, spring
Sika Deer (Cervus nippon) stag in woodland, Arne RSPB Reserve, Dorset, UK, September
Pyrenean merendera / False meadow saffron (Merendera pyrenaica / Colchicum montanum) flowering on montane pastureland, Vegas de Sotres, Picos de Europa, Asturias, Spain, August
Yellow horned poppy (Glaucium flavum) growing on shingle beach, Norfolk, UK, August
Grey seal (Halichoerus grypus) young male swimming above Kelp / Oarweed (Laminaria hyperborea) Farne Islands, Northumberland, UK, North Sea, August
Raft Spider (Dolomedes fimbriatus) female on heathland pool. Note wing of predated damselfly floating on waters surface under the spider. Dorset, UK. August
Harebells (Campanula rotundifolia) growing in disused limestone quarry. Peak District National Park, Derbyshire, UK. August
Stunted oak woodland covered in moss, Wistmans Wood, Devon, UK. August. Highly Commended in the Wild Woods category of the British Wildlife Photography Awards (BWPA) competition 2013
Gannet (Morus bassanus) adult, backlit by the morning sun, in flight. Shetland Islands, Scotland, UK. August
Alpine Ibex (Capra ibex) in its landscape, Valsavarenche, Gran Paradiso national park, Aosta Valley, Italy, May