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Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Ptarmigan (Lagopus mutus) in winter plumage, resting in snow, Svalbard, Norway. April
Willow grouse (Lagopus lagopus) camouflaged against snow, Inari, Kiilopaa, Finland, February
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Red grouse (Lagopus lagopus) on moorland at sunrise, Peak District National Park, UK. August
Red grouse (Lagopus lagopus) in flight over moorland, Yorkshire, UK. September
Black grouse (Tetrao tetrix) males fighting in snow during the breeding season, Viiksimo, Finland. April
Ptarmigan (Lagopus mutus) winter plumage, in flight, Utsjoki, Finland. February
Red grouse (Lagopus lagopus) standing on gritstone rock at sunrise, Peak District National Park, UK. August
Red grouse (Lagopus lagopus) stretching wings at sunrise, Peak District National Park, UK. August
Red grouse (Lagopus lagopus) coming in to land on flowering Heather (Calluna vulgaris), Yorkshire, UK. September
Willow grouse (Lagopus lagopus) winter plumage, in flight, Utsjoki, Finland. February
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
Male Black grouse (Tetrao tetrix) performing courtship display at lek, Cairngorms National Park, Scotland, UK. April
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Male Western capercaillie (Tetrao urogallus) calling during lek mating, Finland. March
Close up of male Western capercaillie (Tetrao urogallus) during lek mating, Finland. March
Black Grouse female (Lyrurus tetrix) perched on frost covered branch, Suomussalmi Finland, January
Black Grouse (Lyrurus tetrix) fighting on icem Finland. March
Hazel Grouse (Tetrastes bonasia), Kuusamo, Finland, December
Red grouse (Lagopus lagopus scoticus) portrait of female, in purple heather, Peak District, UK, September
Red Grouse (Lagopus lagopus scoticus) in purple heather, Peak District, uk
Willow grouse (Lagopus lagopus) sheltering under snow laden tree, Inari Kiilopa Finland January
Willow grouse (Lagopus lagopus) extremely well camouflaged below snow laden tree, Inari Kiilopaa Finland January Highly commended in the Bird category of the Asferico Photography Competition 2017
Red Grouse (Lagopus lagopus scoticus) male on rock in swirling spindrift of snow, Scotland, UK, February. Highly commended in the Habitat category of the BWPA
Willow grouse / ptarmigan (Lagopus lagopus) Agapa River, Taimyr Peninsula, Siberia, Russia
Capercaillie (Tetrao urogallus) male displaying and calling at lek, Kamchatka, Far east Russia, May
Black Grouse (Tetrao tetrix) lek with male displaying and females around in winter, Tver, Russia. April
Black Grouse (Tetrao tetrix) females in tree in winter, Tver, Russia. April
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) males fighting in winter, Tver, Russia. April
Willow grouse / Ptarmigan (Lagopus lagopus) pecking twig, Utsjoki, Finland, April
Spruce grouse (Dendragapus fuliginosus) stops while foraging among fall blueberry bushes. Denali National Park, Alaska, USA
Rock ptarmigan (Lagopus mutus) standing in snow, Myvatn, Thingeyjarsyslur, Iceland, April 2009
Capercaillie (Tetrao urogallus) male calling as part of display, Cairngorms NP, Scotland, February 2009. WWE INDOOR EXHIBITION
RF- Capercaillie (Tetrao urogallus) male displaying in heavy snowfall, Cairngorms National Park, Scotland. February
Willow grouse (Lagopus lagopus) in snow, Dovrefjell National Park, Norway, February 2009
Willow grouse (Lagopus lagopus) on rock in snow, Dovrefjell National Park, Norway, February 2009
Frost on Willow grouse (Lagopus lagopus) feather on Mountain bearberry (Arctous alpinus) covered in frost, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden, September 2008
Male Capercaillie (Tetrao urogallus) displaying in front of camera in a wintry pine forest, Cairngorms National Park, Scotland, February 2009 WWE BOOK, WWE OUTDOOR EXHIBITION
Rock ptarmigan (Lagopus mutus hyperborea) in snow, Spitsbergen, Svalbard, March 2009
Black grouse (Tetrao tetrix) female at lek, Creag Megaidh NNR, Highlands, Scotland, UK, April
Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
Rock ptarmigan (Lagopus mutus) male hidden by rocks, eye visible, winter plumage, Cairngorm Mountains, Cairngorms NP, Highland, Scotland, UK, February
Black grouse (Tetrao tetrix) male in flight at lek, Cairngorms NP, Grampian, Scotland, UK, April
Close up of Capercaillie (Tetrao urogallus) male displaying in pine forest, Cairngorms NP, Highlands, Scotland, UK, February
Rock ptarmigan (Lagopus mutus) perched on boulder, camouflaged against snow in winter plumage, Cairngorms NP, Highlands, Scotland, UK, February
RF- Rock ptarmigan (Lagopus mutus) standing on boulder, camouflaged against snow in winter plumage. Cairngorms National Park, Highlands, Scotland, UK, February
