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Two paint horses, a palomino and a sorrel quarter horse running, Flitner Ranch, Shell, Wyoming, USA
Cavalier King Charles Spaniel puppies aged 7 weeks, with tricolour and blenheim colouration, on hay bale in straw
Four chocolate working cocker spaniels on wall. Mother and offspring. Monmouth, Monmouthshire, Wales
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Peacock butterfly (Inachis io) and Small tortoiseshell butterflies (Aglais urticae) feeding on Buddleia flowers (Buddleia davidii), Derbyshire, UK, September
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
Asiatic lion (Panthera leo persica), two females and two cubs. Gir National Park, Gujarat, India. Photo© Phillip Ross/Felis Images
Four black-and-white Border Collie puppies, age 7 weeks, sitting in a row
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Group of four Giraffes (Giraffa camelopardalis) Mkuze, South Africa
Warthogs (Phacochoerus aethiopicus) mother with babies drinking, Mkhuze game reserve, KwaZuiu-Natal, South Africa, June
Moutain gorilla (Gorilla beringei beringei) close up of hand, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Horsesgrazing on Bulbarrow Hill at dawn, Dorset, England, UK, July 2014
Haflinger horse mares and foals running in meadow. England, UK
Feral domestic rabbits (Oryctolagus cuniculus) sitting up alert, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February
Horses hoof / Tinder fungus (Fomes fomentarius) on tree trunk, European beech (Fagus sylvatica) forest, Stuzica primeval Forest, Unesco World Heritage Site, Poloniny National Park
Mallard (Anas platyrhynchos) ducklings, Bradfield Nature Reserve, Berkshire, England, UK, May
Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Alpaca Herd on Norfolk Farm, winter
Grey-cheeked parakeets (Brotogeris pyrrhoptera) perched and grooming on a branch. Guayaquil, Guayas, Ecuador
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Barn owl (Tyto alba) four chicks in nest, La Pampa, Argentina
Bluebells (Endymion non-scriptus) flowering, Perthshire, Scotland, May
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
Spotted eagle ray (Aetobatus narinari) school, Kagi kandhu, ; North Male Atoll, Maldives, Indian Ocean
Cheetah (Acinonyx jubatus) cubs 4 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
Four Wisent / Bison (Bison bonasus) standing in a row. Bialowieza Forest, Bialowieza National Park, Poland, November
Domestic sheep, lambs playing in a field, Norfolk, UK, March
Small herd of Asiatic Wild Ass (Equus hemionus) running, Gobi National Park, Mongolia
Four British Shorthair kittens, two silver-shaded and two golden-mackerel-tabby, sitting on stand
Pigtail Macaques (Macaca nemestrina) crossing purpose-built primate wires"Pigtail Macaques (Macaca nemestrina) crossing purpose-built " primate wires" connecting isolated patches of forest, Kinabatangan River, Sukau, Sabah, Borneo, September 2008
Rothschild's giraffes (Giraffa camelopardalis rothschildi), feeding on leaves, Woburn Safari Park, UK, June, captive
Great crested grebe (Podiceps cristatus) carrying chicks on its back in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. May
Giant Otters (Pteronura brasiliensis) running along a sand bar. Cuiaba River, Pantanal, Brazil
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Atlantic walruses (Odobenus rosmarus rosmarus) hanging out in shallow water in Svalbard, , Norway, June
Eastern white-bearded wildebeest (Connochaetes taurinus), four running in line, on migration, Masai Mara National Reserve, Kenya
Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK
Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK
Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador
Silhouette of Red deer (Cervus elaphus) stag, hind and fawns at sunset, Norfolk, UK
Hazel nuts (Corylus avellana) Dorset, UK September
Sika deer (Cervus nippon) stags in summer coat, Arne RSPB Reserve, Dorset, England, August
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October