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Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Leopard (Panthera pardus) cub looking up at birds (out of frame) with mother in background, Khwai River, Botswana
Red howler monkey (Alouatta seniculus) howling, captive, digitally modified
Cairns birdwing butterfly (Ornithoptera priamus) male on fern leaf and Blue Mountain Swallowtail butterfly in flight, Atherton Tablelands, Queensland, Australia, October
Skewbald and chestnut Icelandic horses in the snow, Snaefellsnes Peninsula, Iceland, March
Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
RF- Head portrait of a European grey wolf (Canis lupus), with a second in the background, captive, Norway, February
RF - Red Deer stags (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Bottlenose dolphins (Tursiops truncatus) approaching with curiosity, Dolphin Reef, Eilat, Israel. Red Sea
Impala (Aepyceros melampus) female and calves, Masai-Mara Game Reserve, Kenya
Cheetah (Acinonyx jubatus) cubs 4 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
Mountain pansy (Viola lutea) flowers, Strathspey, Cairngorms National Park, Scotland, June
Godrevy Lighthouse at sunset, seen through rocksand rockpool in the foreground. Nr Hayle, Cornwall, UK, June 2009
Domestic sheep, lambs playing in a field, Norfolk, UK, March
Aylesbury ducks following in a line on village green, Weedon, Buckinghamshire, UK, October
Australian Shepherd, red-tri coated, portrait sitting in snow, with picket fence behind
Borage (Borago officinalis) flowers, Menorca, Balearic Islands, Spain, Europe
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Brown-lipped / Grove / Banded snails (Cepaea nemoralis) on plant stem, Menorca, Balearic Islands, Spain, Europe
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
RF - Common blue butterflies (Polyommatus icarus) resting on grass. Vealand Farm, Devon, UK. September
Rock sea-spurrey (Spergularia rupicola) growing on seashore at the base of a cliff, Cornwall, UK, September
Arctic Fox (Vulpes lagopus semenovi), in dark summer pelage, patrolling cliffside rocks along the coast of Medny Island in the Commander Islands (aka Komandorsky Islands), Russia, July
RF- Capercaillie (Tetrao urogallus) male displaying, Inshriach, Cairngorms National Park, Scotland, UK. February
Grasshopper Warbler (Locustella naevia) singing, Joensuu, Finland, May
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Redpoll (Carduelis flammea) male landing on Alder (Alnus glutinosa) Surrey, England. February
Alpine meadow in flower including Scabious flowers. Austrian Alps at 1700 metres altitude, North Tyrol, Austria, June
Lobster pots on harbourside at night, Tobermory harbour, Isle of Mull, Scotland
RF - Early Purple Orchid (Orchis mascula) in flower, Broxwater, Cornwall, UK. May 2016
RF- Pool Frog (Rana lessonae) sitting on white lily pad, Danube delta rewilding area, Romania
Chilean huemul or South Andean deer (Hippocamelus bisulcus), Torres del Paine National Park, Patagonia, Chile. Endangered species
Marine Iguanas (Amblyrhynchus cristatus) on the south coast of Espanola Island, Galapagos, Vulnerable species
Domestic pig, hybrid large white sow in free-range sty, UK, August 2010
Dalmation dogs playing on beach, Norfolk, UK, August
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Wild gladiolus (Gladiolus illyricus) flowers, Menorca, Balearic Islands, Spain, Europe
Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India
RF - Harts tongue fern (Phyllitis scolopendrium) leaf unfurling and creating the shape of a heart, Cornwall, UK, May
Cane Toad / Marine Toad / Giant Toad (Bufo marinus) adult jumping in Dogweed (Dyssodia pentachaeta) field. Laredo, Webb County, South Texas, USA, April
RF - Andean condor (Vultur gryphus) flying over Torres del Paine Massif. Torres del Paine National Park, Chilean Patagonia, Chile. March
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
RF- Red Admiral butterfly (Vanessa atalanta) perched on Bigtooth Maple (Acer grandidentatum). Lost Maples State Park, Hill Country, Central Texas, USA. November
