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Female Harpy Eagle (Harpia harpyja) perched close to its nest. Pousada Currupira d Araras, south west Brazil
German Shepherd Dog bitch pup, Coco, 14 weeks old, with raised paw
Regal Jumping Spider (Phidippus regius) female. Captive, endemic to North America
Grey wolf (Canis lupus) mother and two month old cubs, pups begging for food by licking mothers mouth, captive
Marsh harrier (Circus aeruginosus) adult female circling over Oilseed rape crop, Norfolk, England, May. Digital composite
Beagle, bitch resting on hay bale in straw
On a misty morning, female Grizzly bear (Ursus arctos horribilis) standing up in alert and looking around with her cubs for danger, while feeding on sedges, Khutzeymateen Grizzly Bear Sanctuary
Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
Brown bear (Ursus arctos) and cubs in the snow, Finland, April
Coastal brown bear (Ursus arctos) female in a meadow, cubs nearby, Lake Clarke National Park, Alaska, USA, July
Rio pescado harlequin toad (Atelopus balios) female on branch, Azuay, Ecuador, Critically Endangered, possibly extinct
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Snowy owl (Bubo scandiaca) close up of female landing on snow covered ground, Canada, February. Magic Moments book plate
Bateleur Eagle (Terathopius ecaudatus) female, stretching wings Kgalagadi Transfrontier Park, South Africa. January
Four-spot orb weaver spider (Araneus quadratus) male approaching larger female to mate. Peak District National Park, Derbyshire, UK. October
Female chalkhill blue butterfly (Lysandra coridon) with wings closed resting on Devils-bit scabious (Succisa pratensis), Hatch Hill, Somerset, UK. August 2016
Raft Spider (Dolomedes fimbriatus) female on heathland pool. Note wing of predated damselfly floating on waters surface under the spider. Dorset, UK. August
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Red deer (Cervus elaphus) female amongst autumn foliage, Arran, Scotland, October 2013
Shelduck (Tadorna tadorna) female with ducklings, Cley, Norfolk, East Anglia, England, UK, July
Female Kestrel (Falco tinnunculus) perched in the entrance to a nest hole in an old tree, Hertfordshire, England, UK, June
Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Common blue butterfly (Polyommatus icarus) female, Southwest London, England, UK. June
Female Ibex (Capra ibex) portrait, Creux du Van Natural Reserve, Val de Travers, Switzerland, December
Cheetah (Acinonyx jubatus) female with cubs sitting in long grass during the dry season, Masai-Mara Game Reserve, Kenya. March
Female Tassled wobbegong shark (Eucrossorhinus dasypogon) rests alongside stand of Barrel sponges (Xestospongia testudinaria) Blue Magic, Raja Ampat, West Papua, Indonesia
Olive Ridley Sea Turtle (Lepidochelys olivacea) nesting, Arribada (mass nesting event), Playa Morro Ayuta, Oaxaca state, southern Mexico. Vulnerable species
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
Gloucester old spot piglets (Sus scrofa domestica) suckling at the Cotswold Farm Show, Cirencester, UK, July 2008
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Red Fox (Vulpes vulpes) vixen on a misty day in woodland, Black Forest, Germany, July. Runner-up in Animal Portraits category, Wildlife Photographer of the Year competition 2011
Forester kangaroo (Macropus giganteus tasmaniensis) family group, male, female and large joey, Tasmania, Australia, February. Subspecies of Eastern grey kangaroo
Black Labrador and two puppies
Andalusian mare running in snow storm, Colorado, USA
Red fox (Vulpes vulpes) vixen yawning in the morning light (captive). Lifton, Devon, UK, May
Domestic pig, hybrid large white sow in free-range sty, UK, August 2010
Domestic pig, free range sow and piglet sniffing noses, UK, August 2010
Great Dane, black coated bitch, and Magyar Vizsla / Hungarian Pointer, tan coated maleGreat Dane, black coated bitch, and Magyar Vizsla /Hungarian Pointer, tan coated male, double portrait in profile together
Roan antelope (Hippotragus equinus) with young offspring, Mlilwane nature reserve breeding programme, Swaziland, Southern Africa
Polar bear (Ursus maritimus) sow with her cub outside their den in late winter, Arctic coast of Alaska
Maine Coon female cat, Serafin, in fierce defensive posture
Border Terrier bitch puppy, Kes, with head cocked on one side
Red leaf monkey (Presbytis rubicunda) mother and baby, Danum Valley, Sabah, Borneo, Malaysia August
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
Yellow Labrador Retriever bitch pup, 10 weeks, with a red rose and crossed paws
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana