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Ghost glass frog (Centrolenella ilex) portrait, Costa Rica
Portrait of a young Little Owl (Athene noctua)
Arctic fox (Vulpes lagopus), juvenile looking at camera, portrait, winter pelage. Dovrefjell National Park, Norway. September
Panther chameleon (Furcifer pardalis) sitting on branch with black background, Nosy Faly, Madagascar
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Goldenrod crab spider (Misumena vatia) yellow female on Honesty flower, Bristol, UK, April
Arctic fox (Vulpes lagopus) portrait of juvenile, winter pelage. Dovrefjell National Park, Norway. September
Bornean horned frog (Megophrys nasuta) on forest floor. Danum Valley, Sabah, Borneo, Malaysia
Imbabura tree frog (Hypsiboas picturatus) portrait, Canande, Esmeraldas, Ecuador
Female Scottish wildcat (Felis silvestris grampia) portrait, in winter coat, semi habituated living wild in native oakwood, Lochaber, Scotland, April
RF - White-legged damselfly (Platycnemis pennipes) peering over Oxeye daisy (Leucanthemum vulgare) flower. The Netherlands. August
Portrait of a Common Wasp (Vespula vulgaris). UK. Focus stacked image (dead specimen)
Aye-aye (Daubentonia madagascariensis) extracting beetle grubs from wood. Endemic to Madagascar. Photographed under controlled conditions at Durrell Wildlife Conservation Trust, Jersey, UK. captive
Turbot (Scophthalmus maximus) detail of the eyes and mouth, United Kingdom, September
Red eyed tree frog (Agalychnis callidryas) portrait Captive
Blue eyed tabby and white Siberian cross kitten, age 13 weeks
Close up of Garibaldi fish face {Hypsypops rubicunda}, Channel Islands, California, USA
Butterfly / Rhinoceros viper (Bitis nasicornis). Captive. Occurs in West and Central Africa
Dragon mantis (Toxodera beieri), detail of head showing large compound eyes giving it the visual acuity to hunt fast-moving prey. Danum Valley, Sabah, Borneo
Deep sea fish (Winteria telescopa) with forward pointing tubular eyes
Smooth knob-tailed gecko (Nephrurus levis occidentalis) portrait. Kalbarri, Kalbarri National Park, Western Australia. October
Cuttlefish (Sepia officinalis) close up of eye, Tenerife, Canary Islands
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Bubble eye goldfish (Carassius auratus) with upward pointing eyes and two large fluid-filled sacs.. In captivity, Shanghai Zoo, Shanghai, China
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Caribbean reef squid (Sepioteuthis sepioidea) swimming in open water at night, Guadeloupe Island, Mexico. Caribbean
Crocodile monitor (Varanus salvadorii) close up eye, captive, occurs in New Guinea
Close up if a Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Bullseye / Io moth (Automeris io) showing eye spot markings on wings during deimatic display to deter predators, photographed on a white background, originating from North and Central America
Indian moon / Indian luna moth (Actias selene) head-on view showing feather-like antennae. Captive
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Sumatran tiger (Panthera tigris tigris / Panthera tigris altaica) close up portrait, captive
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
Wild boar (Sus scrofa) eyes glowing at dusk, introduced species in La Pampa, Argentina
South Georgia pintail (Anas georgica georgica) swimming in front of Southern elephant seal (Mirounga leonina), male. Gold Harbour, South Georgia. October
Grey whale (Eschrichtius robustus) eye, Magdalena Bay, Baja California, Mexico, February. Winner in the Ocean Views category of the 2018 Natures Best Winland Smith Rice Awards
Lava lizard (Microlophus peruvianus) juvenile peering out of eye of dead sea lion. Paracas National Reserve, Peru. Highly commended in the Animals in their Environment Category of Wildlife
Small emperor moth (Saturnia pavonia) on Speedwell flowers, Annagarriff Wood NNR, County Armagh, Northern Ireland, UK, May
Red-eyed tree frog {Agalychnis callidryas} resting in Bromeliad flower, Nicaragua, June
Jumping spider (Salticidae sp), close up. Wuliangshan Nature Reserve, Jingdong, Yunnan Province, China
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
Wild female Amur leopard (Panthera pardus orientalis) looking up, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Male Lion (Panthera leo) portrait, close-up of face, captive, occurs in Africa
Portrait of a Birman-cross cat
Fillys eye, wild Mustang horse at Black Hills Wild Horse Sanctuary, South Dakota, USA
Devil scorpionfish (Scorpaenopsis diabolus) close up of eye, Pak Lap Tsai, Sai Kung, Hong Kong, China
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017