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Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July
Aggregation of thousands of moulting Spider crabs (Leptomithrax gaimardii), South Australia Basin, Australia. Pacific Ocean
Sally lightfoot crabs (Grapsus grapsus) on rock with waves, Cape Douglas, Fernandina Island, Galapagos
Pederson cleaner shrimp (Periclimenes pedersoni), Cienaga de Zapata National Park, Matanzas Province, Cuba
Arrow crab (Sternorhynchus lanceolatus). Tenerife, Canary Islands
Commensal emperor shrimp (Periclimenes imperator) hitching a ride on a Nudibranch (Risbecia tryoni). Lembeh Strait, North Sulawesi, Indonesia
Anemone prawn (Periclimenes sagittifer) on Snakelocks anemone (Anemonia viridis) Jersey, British Channel Islands, June
Whip coral shrimp (Pontonides uncigar) Bluff Island, south of Sai Kung Peninsula, Hong Kong, China
Land crab (Geosesarma sp. ) which raids Pitcher plant (Nepenthes ampullaria) for preyLand crab (Geosesarma sp.) which raids Pitcher plant (Nepenthes ampullaria) for prey, Sarawak, Borneo
Aggregation of thousands of Spider crabs (Leptomithrax gaimardii) for moulting, South Australia Basin, Australia. Pacific Ocean
European / Common Lobster (Homarus gammarus) in rocky retreat. Controlled conditions, Cornwall, UK. May
Spiny spider crab (Achaeus spinosus) reaching up and scratching its head, while climbing on purple soft coral bush (Dendronephthya sp.) Sinai, Egypt. Red Sea
Marine iguana (Amblyrhynchus cristatus) and Sally-lightfoot crabs (Grapsus grapsus) on rocks at high-tide with sea spray, Galapagos Islands, South America
Mount Lewis spiny crayfish (Euastacus fleckeri) in defensive posture with claw raised, at edge of mountain stream, Mount. Lewis, Wet Tropics World Heritage area, Queensland, Australia. Endangered
Mexican Fiddler Crab (Leptuca crenulata) male displaying, Morua Estuary, near Rocky Point, Gulf of California, Mexico, June
Hermit crab (Dardanus calidus) Vis Island, Croatia, Adriatic Sea, Mediterranean
Lamington spiny crayfish (Euastacus sulcatus) blue form, in leaf litter under tree roots with claws raised, rare crayfish that commonly forages away from water on the forest floor above 300m
Galapagos sea lion (Zalophus wollebaeki) inquisitive pup playing with Sally Lightfoot Crab (Grapsus grapsus). Plazas Island, Galapagos, Ecuador
Swimming crab (Cronius ruber) Canary Islands, Tenerife. Invasive species
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
Strawberry land hermit crab (Coenobita perlatus). On white coral sand beach, Small unhinhabited coral islet in Raa Atoll lagoon, Maldives
Columbus crab (Planes major) which was stranded on the beach, South Africa
Boxer Crab (Lybia tessellata) with sea anemones in its claws for defense. Lembeh Strait, North Sulawesi, Indonesia
Hermit crab (Pagurus bernhardus) on yellow sponge, Loch Etive, west coast of Scotland, UK, September
Horse mussel (Modiolus modiolus) bed with Brittlestars (Ophiothrix fragilis) and Edible crab (Cancer pagurus), Shetland, Scotland, UK, September
Common lobster (Homarus gammarus) looks out from its hole amongst Dead mans fingers (Alcyonium digitatum) beneath a forest of Oarweed kelp (Laminaria hyperborea)
Princely fiddler crab (Uca princeps), Bahia Magdalena, Baja California Peninsula, Mexico, June
Marbled swimming crab (Liocarcinus marmoreus) Jersey, British Channel Islands, June
Common Hermit Crab (Pagurus bernhardus) Bouley Bay, Jersey, British Channel Islands
Hairy Crab (Pilumnus hirtellus) on seaweed on the beach, Sark, British Channel Islands
Emperor shrimp (Periclemenes imperator) on a nudibranch (Ceratosoma sp. ) Lembeh StraitEmperor shrimp (Periclemenes imperator) on a nudibranch (Ceratosoma sp.) Lembeh Strait, North Sulawesi, Indonesia
Fiddler crab (Uca) male, hiding in an old shell at Ha Pak Nai mudflat, Yuen Long District, Hong Kong, China
American / Northern lobster (Homarus americanus) partially hidden under seaweed, Nova Scotia, Canada, July
White forest crab (Cardisoma armatum) portrait, seen through claw, Island of Principe UNESCO Biosphere Reserve, Democratic Republic of Sao Tome and Principe, Gulf of Guinea
Harlequin shrimp (Hymenocera elegans) on blue sea star. This species of shrimps feeds on starfish, turning them upsidedown and keeping them as living larder. Bitung, North Sulawesi, Indonesia
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
American oystercatcher (Haematopus palliatus) chasing after Sally lightfoot crab (Grapsus grapsus) across shoreline. Galapagos Islands, Ecuador
Yellow fiddler crab (Uca sp. ) portrait, Sunderban tiger reserve, West Bengal, IndiaYellow fiddler crab (Uca sp.) portrait, Sunderban tiger reserve, West Bengal, India
Red rock crab (Grapsus adscensionis) grazing on algae from rock, Tenerife, Canary Islands, October
Sally Lightfoot crabs (Grapsus grapsus) on rocks caught in the swell, Espanola Island, Galapagos National Park, Galapagos Islands
Sally Lightfoot crab (Grapsus grapsus) on beach, portrait, Santiago Island, Galapagos National Park, Galapagos Islands
Deep red Feather star shrimp (Hippolyte prideauxiana) crawling secretly amongst the arms of a Feather star (Antedon bifida), Loch Duich, Highands, Scotland, UK
Pair of Coleman shrimps (Periclimenes colemani) living in a fire urchin (Asthenosoma varium). The female is the larger shrimp
Bullseye reef lobster (Hoplometopus holthuisi) on rocky seabed, Hawaii, Pacific Ocean
Harlequin Shrimp (Hymenocera elegans) on sea floor. Huatulco Bays National Park, Oaxaca state, Mexico, Pacific Ocean. November
Velvet swimming crab (Necora puber) male, in defensive posture, with wreck of the Rosalie in background, Weybourne, Norfolk, England, UK, North Sea
